New or poorly known species of Sinodrepanus Simonis, 1985 (Coleoptera: Scarabaeidae: Oniticellini), with the first identification key for the genus Author Montanaro, Giulio text Zootaxa 2024 2024-11-12 5537 4 577 595 http://dx.doi.org/10.11646/zootaxa.5537.4.10 journal article 10.11646/zootaxa.5537.4.10 1175-5326 14240160 4BDF6685-879A-474B-BD8D-FC23323A0947 Sinodrepanus falsus ( Sharp, 1875 ) Oniticellus falsus Sharp, 1875: 52 (original description) Drepanochirus falsus : Boucomont & Gillet, 1921: 19 Cyptochirus falsus : Boucomont & Gillet, 1927: 103 ; Paulian, 1945: 128 Drepanocerus falsus : Janssens, 1953: 36 , 37; Balthasar, 1963: 62 , 66; Biswas & Chatterjee, 1985: 151 (probably to be ascribed to S. schoenfeldi new species ); Sewak, 2003: 253 , 283 (to be ascribed to S. schoenfeldi new species ?); Sewak, 2006: 219 Sinodrepanus falsus : Simonis, 1985: 98 , 103, 104; Kabakov & Napolov, 1999: 73 ; Ochi et al ., 2004: 355 ; Bezděk & Krell, 2006: 157; Krikken, 2009: 9 , 14, 21; Kabakov & Shokhin, 2014: 58 ; Mittal & Jain, 2015: 393 ; Bezděk, 2016: 174 ; Montanaro, 2021: 585 Type locality : Laos. Type material examined . Holotype , ♀ ( MNHN ): Laos / Mouhot [wc, line 1 hb, line 2 pb] // Oniticellus / falsus / Type D.S. [wc, hb except “Type” pb] // Ex Musaeo / D. Sharp 1890 [wc, pb] // MUSÉUM PARIS / 1952 / COLL. R. OBERTHÜR [wc, pb] // A. Janssens det. 1953 / Drepanocerus / falsus Sharp ♀ / Type [wc, hb except “A. Janssens det. 195” pb] // TYPE [rc, pb] // HOLOTYPE [rc, pb] // HOLOTYPE / Sinodrepanus / falsus (Sharp) [wc, pb] // MNHN / EC8103 [wc, pb] . Additional material examined ( 37♂♂ , 20♀♀ ). China : S Yunnan province , Xishuangbanna , Guangpin environment, 0km [sic!] N Jinghong , 1000m , 19–27.i.2003 , S. Murzin legit [ 1♂ , PMOC ] ; S Yunnan province , Jinghong , 9–14.vii.1990 , R. Červenka legit [ 2♂♂ , NMPC ] . Laos : Xayaboury province , Hongsa environment, 19°40–44’N 101°20’E, 550–750m , 2–3.vii.2010 , D. Hauck legit [ 1♀ , JSCS ] ; Paklay , viii.1917 , R.V. de Salvaza legit [ 1♂ , BMNH ] ; Louangphrabang , Thong Khan , 19°35’N 101°58’E , ~ 750m , 11–21.v.2002 , Vit Kubáň legit [ 10♂♂ & 6♀♀ , NMPC ] ; Phongsaly province , Ban Sano Mai environment, 21°21’N 102°03’E , ~ 1150m , 19–26.v.2004 , Vit Kubáň legit [ 2♂♂ , NMPC ] ; Xayaboury province , Pak Lay env., 18°13’N 101°22–24’E, 12–13.vii.2010 , David Hauck legit [ 4♂♂ , 2♀♀ , NMBS ; 2♂♂ , 1♀ , JSCS ] . Thailand : Ban Tao Phan, 11.iii.1918 [likely Ban Phan Tao in Chiang Mai province ; 1♂ , BMNH ] ; Chiang Mai province , Chiang Dao , Chiang Dao Hill Resort environment, 19°33’28.37”N 99°4’34.20”E , 2–11.vii.2019 , Vishnyakov Alexey legit [ 1♂ , GMOC ] ; Chiang Mai province , Chiang Dao , 19°33’28”N 99°4’34”E , 14.vi.2017 , Prosvirov legit [ 1♀ , GMOC ] ; Chiang Mai province , Chiangdao Hill Resort , 19°33’29.19”N 99°04’33.34”E , 1-15.vii.2019 , Ustinov V. legit [ 2♂♂ , GMOC ] ; 100km N Chiang Mai , Chiang Dao hill Resort , 20-30.VI.2008 , S. Murzin legit [ 1♂ , PMOC ] ; Loei province , Na Haeo biological station , cow dung , 5–12.v.2001 , Constant & Grootaert legit [ 4♂♂ , 6♀♀ , RBINS ] ; Loei province , Na Haeo , zebu dung , 18.v.2003 , J. Constant & K. Smets legit [ 2♂♂ , 2♀♀ , RBINS ] ; Umphang river , 16°07’N 99°00’E , 1000m , 28.iv– 6.v.1991 , Vit Kubáň legit [ 4♂♂ , 1♀ , NMPC ] . Diagnosis . Sinodrepanus falsus can be distinguished from other species by the characters provided by Montanaro (2021) . Additionally, this species is rather distinctive for having a particularly uneven body surface, with strong pronotal carinas that delimit well-marked spaces on the pronotum, recalling those of the genus Cyptochirus Lesne, 1900 —see also Simonis (1985) and Simonis & Zunino (1980) . See also the dichotomous key provided below. Description : see Montanaro (2021) . Variation . Body size ranges from 7.2 to 9.8 mm . No other significant variation was observed. Verified distribution . China ( Yunnan ), Laos , Thailand ( Fig. 4 ). Kabakov & Napolov (1999) recorded S. falsus from Vietnam , considering S. exsul as a synonym of it. Kabakov & Shokhin (2014) recorded S. falsus from China ( Sichuan ). These records should be reassessed to understand the true identity of such species. See also the “ Indian Sinodrepanus : an overlooked diversity?” section for more distributional comments. Ecology . Some specimens were collected in cow and zebu dung. Etymology . Latin adjective meaning “deceived”, “tricked”; by extension “false”, “untrue”. It is probably referred to the cryptic appearance of this species, shared with all other Sinodrepanus , that camouflages these beetles within dung and other fibrous substrates.