Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443 Author Soto, Eduardo M. Author Ramírez, Martín J. text Zootaxa 2012 2012-08-29 3443 1 1 65 https://biotaxa.org/Zootaxa/article/view/zootaxa.3443.1.1 journal article 10.11646/zootaxa.3443.1.1 1175­5334 5254614 Philisca doilu (Ramírez) Figs 6–8 , 34e , 38e Liparotoma doilu Ramírez, 1993 (female holotype , from Argentina : Neuquén : Lanín Natl. Park: Tromen Lake , I.1991 , V. Fiorani , MACN, reexamined). Philisca doilu : Ramírez 2003: 191 . Diagnosis: Resembles P. hyadesi by the low cephalic area ( Ramírez 1993 : fig. 30), the transversely wrinkled median field of the epigyne advanced from the epigastric furrow ( Fig. 6b ), and the elongate paramedian apophysis ( Fig. 8 ). It can be distinguished by the narrower sternum ( Fig. 7c ; width less than 63% of the sternum length). Description: Female : described in Ramírez (1993) . FIGURE 6. Female of Philisca doilu Ramírez (MACN-Ar 26762). ( a female dorsal habitus; b epigyne, ventral view; c cleared epigyne, dorsal view). (AB: accessory bulbs, CD: copulatory ducts, MF: median field, S: spermathecae). FIGURE 7. Philisca doilu Ramírez (MACN-Ar 15097). a–d male ( a dorsal habitus; b prosoma, dorsal view; c ventral habitus; d prosoma, ventral view). FIGURE 8. Philisca doilu Ramírez (MACN-Ar 15097). Male, a–c left palp ( a prolateral view, b ventral view, c retrolateral view); d–h copulatory bulb ( d prolateral view, e ventral view, f retrolateral view, g apical view, h apical–retrolateral detail view). (C: conductor, E: embolus, MA: median apophysis, PMA: paramedian apophysis, rPMA: retrolateral cusp of the paramedian apophysis). Male (MACN-Ar 15097): Total length 11.97. Prosoma length 5.45, width 4.26. Length of tibia/metatarsus: I , 6.12/4.79; II , 5.05/4.26; III , 2.39/2.53; IV , 3.72/3.46. Chelicerae very strong ( Fig. 34e ), with two teeth on retromargin (basal tooth larger) and three on promargin (apical one very large). Sternum length 2.93, width 1.87. Spines: leg I , femur d 1–1–1, p 2ap. II , femur d 1–1–1, p d2ap. III , femur d 1–1–1, p and r d1ap or 0; patella r d1; tibia v 1–2ap or 0–2ap, p 0–1, r d1–1; metatarsus v 2–0–2, p and r d1–2ap. IV , femur d 3–2–1 or 2–2–1, r d1ap or 0; patella r d1; tibia v 2ap or p1ap, r d1–1; metatarsus v 2–p1–p1–2 or 2–0–0–2, d 2ap, p 1ap, r d1–1. Opisthosoma length 6.38, spiracle–epigastrium 2.80, spiracle–spinnerets 0.53. Color in ethanol ( Fig. 7 ): Prosoma and chelicerae reddish brown with white pubescence. Sternum brown. Legs brown with black spots, leg I darker than the others. Opistosoma light brown, with dorsal and ventral brown pattern. Femur IV in dorsal view with 2–4 spines at base. Palp ( Fig. 8 ): Tibia width/length 0.28. Embolus short, anterior margin longitudinally flattened, slightly projecting in a ridge; basal process flattened, rounded ( Fig. 8 ). MA long, with one big branch at base ( Fig. 8h ). PMA long and sinuous. Sperm duct without apical or retrolateral loop. pPMA simple, wide. Conductor complex, with canal, dentate prolateral ridge absent, prolateral portion with triangular, flattened projection; retrolateral portion thin, projecting in acute peak. Variability: Spines: Females: III , tibia v 0–p1–2; IV , femur d 3–1–1, p 0, r d1ap; tibia r d1–d1–0–1. Males: IV , femur d 4–1–1 or 3–1–1, p d1ap. Distribution: Southern forests in Chile , from Arauco (Region VIII) to Valdivia provinces (Region XIV), and in Argentina , in Neuquén province ( Fig. 38e ). New records: ARGENTINA : Neuquén : Cerro La Mona , S40,5678º W71,69727º , 1500m , I.2006 , V. Werenkraut , Pitfall , 1 male ( MACN-Ar 15097 ) , 2 males ( MACN-Ar 15098 ) . CHILE : Región IX ( Araucanía ): Malleco Prov. : Nahuelbuta Natl. Park , S37º47’ W73º00’ , 1200m , 12.II.2005 , J. Barriga T., fogging Nothofagus dombeyi , 1 female ( MACN-Ar 26762 ) , 1 immature ( MACN-Ar 26761 ) ; Malalcahuello Way Lonquimay , fogging, 1236m , 15ºC S38º26.377’ W71º30.540’ , N. dombeyi , 35m , 9:47AM, 19.I.2007 , Arias et al. Berkeley , 1 immature ( CASENT 9039004 ) ; Malalcahuello Way Lonquimay , fogging, 1221m , 29°C, S38°26.388’ W71°30.613’ , N. dombeyi , 50m , 6:55PM, 18.I.2007 , Arias et al. Berkeley , 1 immature ( CASENT 9039011 ) ; Malalcahuello Way Lonquimay , fogging, 1236m , 15ºC, S38º26.377’ W71º30.540’ , N. dombeyi 35m , 9:47AM, 19.I.2007 , Arias et al. Berkeley , 3 immatures ( CASENT 9039005 ) . Región XIV (de los Ríos ): Valdivia Prov. : Valdivia , Chaihuin , Reserva Costera Valdiviana , Las Garzas , S39°59.640’ W73°35.197’ , 360m , 20.9° C Fogging 150cc/l Nothofagus nitida , 11:00AM, 27.II.2008 , Arias et al. UCB, 1 immature ( CASENT 9039015 ) ; Valdivia , Chaihuin , Reserva Costera Valdiviana , Las Garzas , S39°49.687’ W73°15.227 , 360m, 17.3°C, fogging 150cc/l N. nitida , 11:00AM, 27.II.2008 , Arias et al., 1 female and 9 immatures ( CASENT 9039063 ) , 1 female ( CASENT 9039067 ) . Material re-examined: ARGENTINA : Neuquén : Lanín Natl. Park, Lago Huechulaufquen , Coloradas , 1.I.1985 , M. Ramírez , 1 female ( MACN-Ar 14800 ) .