Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica Author Fleming, AJ Author Wood, D. Monty Author Smith, M. Alex Author Hallwachs, Winnie Author Janzen, Daniel H text Biodiversity Data Journal 2018 6 11223 11223 http://dx.doi.org/10.3897/BDJ.6.e11223 journal article http://dx.doi.org/10.3897/BDJ.6.e11223 1314-2828-6-11223 Calolydella aureofacies Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029582 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Pereira ; individualID: DHJPAR0029582; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 08-SRNP-36592, ASHYM1003-09; Taxon: scientificName: Calolydellaaureofacies; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: aureofacies; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Ponderosa; verbatimElevation: 1060 ; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Cyclophora Janzen23 ; verbatimEventDate: 26-Sep-2008 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029601 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Pereira ; individualID: DHJPAR0029601; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 08-SRNP-36589, BOLD:AAH9540, ASHYM1022-09; Taxon: scientificName: Calolydellaaureofacies; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: aureofacies; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Ponderosa; verbatimElevation: 1060 ; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Cyclophora Janzen23 ; verbatimEventDate: 26-Sep-2008 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Female: 6mm (Fig. 5). Head (Fig. 5b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose along upper half; parafacial almost entirely gold (>90% coverage); palps black basally. Thorax (Fig. 5a, c): gold on both dorsal and lateral surfaces; four regular thoracic vittae; postpronotum with three setae; 2:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae absent. Wing vein R4+5 with 5-7 setulae dorsally, extending almost to crossvein R-M. Abdomen (Fig. 5a): ground color black, typically with uninterrupted transverse marginal pollinose bands (these can sometimes appear interrupted by a thick median stripe under certain angles of light); both dorsal and ventral surfaces concolorous, silver; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with a pair of discal setae. Male: not known at this time. Diagnosis Calolydella aureofacies can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, female palps black basally, anatergite with three or more hair-like setae, and scutellar discal setae absent. Etymology The specific epithet is derived from the Latin adjective "aurum", meaning gold, and "facies", meaning face, in reference to its gold parafacial and fronto-orbital plate. Distribution Costa Rica, ACG, Guanacaste Province, Sendero Ponderosa, 1060m. Ecology Calolydella aureofacies has been reared once from Cyclophora Janzen23 ( Lepidoptera : Geometridae ), in cloud forest.