Typification of eight current and seven related names and a new section in the genus Bromus (Bromeae, Pooideae, Poaceae)
Author
Llamas, Felix
Author
Acedo, Carmen
text
PhytoKeys
2019
121
53
72
http://dx.doi.org/10.3897/phytokeys.121.30254
journal article
http://dx.doi.org/10.3897/phytokeys.121.30254
1314-2003-121-53
A90CEB1FFFFDE325FFC1FFD53E14FF93
2702029
Bromus lepidus Holmb., Bot. Not. 1924: 326 (1924)
Bromus lepidus
Holmb., Bot. Not. 1924: 326 (1924).Type Protologue: "Nach ROUY kommt sie in Frankreich hier und da vor,aber ziemlich selten;
Kroesche
hat sie nur von einem
Standort
in Norddeutschland. In Schweden kommt sie besonders in Schonen vor; ich habe sie mehrmals eingesammelt, wie in der Gegend von Lund und
Malmoe
an mehreren Orten;
Svenshoeg
in
Wallkaerra
; Skartofta in
Oeved
; Gudmundtorp.
Ausserdem
sah ich Exemplare aus
Blommeroed
in
Oeved
(leg. P. Boren 1903), Skelderviken (leg. Sten Selander), Kalmar (leg. N. Blomgren),
Boras
(leg. C. Sandberg), Fyen, Stenlose (leg. G. Samuelsson). Kommt in Klee- und Grasfeldern, auf Rainen,
Wegraendern
etc. vor, oft mit
B. mollis
und
B. commutatus
zusammen; bei
Malmoe
auch als Ruderat". Type: SWEDEN. Scania:
Malmoe
, in ruderatis, 18-06-1920, Otto R.Holmberg, (lectotype, designated here: LD [LD1136595 image!]; isolectotype: K [K000913599!]). (Figure 7)
Bromus
lepidus Holmb. f. lasiolepis
Holmb., Bot. Not. 1924: 326 (1924). Type Protologue: not indicated. Type: Sweden. Scania:
Vallkoerra
.
Svenenshoeg
. 4-07-1923. Otto R.Holmberg, (lectotype, designated here: LD [LD 1136235 image!]; isolectotype: K [K000913598!]). (Figure 8)
Remarks.
This taxon has a long history full of nomenclatural problems.
Duval-Jouve (1865
: 208), who only lists, without naming them, the variations in spikelet size and hairiness of
Bromus
species, e.g.
Bromus mollis
"
microstachys
glabre" and
Bromus mollis
"
microstachys
pubescent". This is the first mention of this taxon. Later
Rouy (1913
: 236) proposed a named variety under the genus
Serrafalcus
Parl.:
S. mollis
ss
microstachys
Rouy, giving "
Bromus microstachys
Duval-Jouve" (1865
: 207) as a synonym and adding a diagnosis. Therefore, there is no doubt this is the same plant cited by Duval-Jouve (l. c.). Afterwards,
Kroesche
(1924
: 329) described
B. gracilis
Kroesche
, which unfortunately is a posterior homonym as
Weigel (1772
: 15) previously proposed this name for a different plant.
Finally,
Holmberg (1924
: 326) solves this unfortunate situation and names simultaneously the new taxa
Bromus lepidus
Holmb. and
B. lepidus f. lasiolepis
Holmb. The herbaria having Holmberg material are K, LD and S. There are 13 sheets in LD collected before 1924, five in S and three in K. It is reasonably certain that Holmberg studied all those plants before the description of the species and form and that all of them are original material. Therefore, we decided to limit our choices to sheets with the annotations "
B. lepidus
mihi" and "
B. lepidus f. lasiolepis
Holmb.", as this annotation indicates that Holmberg is interpreting them as the new taxa he is going to describe.
Figure 7.
Lectotype of
Bromus lepidus
Holmberg, preserved at the Herbarium of the biological museum of the Lund University (LD1136295), reproduced with permission.
Figure 8.
Lectotype of
Bromus lepidus f. lasiolepis
Holmberg, preserved at the Herbarium of the Biological Museum of Lund University (LD 1136235). Reproduced with permission.