A synopsis of the exindusiate species of Cyathea (Cyatheaceae-Polypodiopsida) with bipinnate-pinnatifid or more complex fronds, with a revision of the C. lasiosora complex Author Lehnert, Marcus text Phytotaxa 2016 2016-01-08 243 1 1 53 http://dx.doi.org/10.11646/phytotaxa.243.1.1 journal article 10.11646/phytotaxa.243.1.1 1179-3163 13682214 65. Cyathea tortuosa Moran (1991: 99) . Type:— ECUADOR . Napo : Reserva Biológica Jatún Sacha, ca. 8 km ESE de Misahualli, 01º04’S , 77º36’W , 450 m , R.C. Moran & C. Rohrbach 5187 ( holotype MO!, isotypes DUKE-10000940!, F-0075659F–0075666F!, QCNE-174–178!, UC-1586249–1586251!, US-00386073–00386075!) Distribution and habitat:Colombia , Venezuela , Ecuador and Peru at 300–1200(–1600) m in humid lowland and premontane forests.