Eleven new species of Amblypsilopus Bigot (Diptera: Dolichopodidae: Sciapodinae) and a key to the species of Madagascar and adjacent islands Author Grichanov, Igor Ya. text European Journal of Taxonomy 2021 2021-06-18 755 1 47 87 http://dx.doi.org/10.5852/ejt.2021.755.1399 journal article 6004 10.5852/ejt.2021.755.1399 d7d8ccff-7a58-4fc8-9eec-c2588b457d56 2118-9773 4983303 4C7C9B68-4026-45F9-98C4-430FE018119F Amblypsilopus flavus ( Vanschuytbroeck, 1962 ) Fig. 3 Megistostylus flavus Vanschuytbroeck, 1962: 353 . Type locality: Madagascar : Ambodivoangy. Chrysosoma flavum – Dyte & Smith 1980: 445 . Plagiozopelma flavum – Bickel 1994: 231 . Amblypsilopus flavus – Grichanov 1998: 84 . Diagnosis Amblypsilopus flavus is very close to A. friedmani sp. nov. in habitus, differing from the latter species in the fore and mid femora with at most few elongate anteroventral and posteroventral setae, at most half as long as femur diameter; the fore tibia and basitarsus without groups of elongate ventral setae at base; the cercus with one bristle between lobes, half as long as ventral lobe. Material examined Holotype MADAGASCAR • ♂ ; Ambodivoangy ; Dec. 1949 ; I. Vadon leg.; P. Vanschuytbroeck det. 1961?; Megistostylus flavus n. sp. ; [ex.] Coll. Mus. Congo; MNHN . Paratype MADAGASCAR • 1 ♂ ; Ambodivoangy ; Dec. 1949 ; I. Vadon leg.; P. Vanschuytbroeck det. 1961?; Megistostylus flavus n. sp. ; Coll. Mus. Congo; MRCA . Additional material MADAGASCAR • 1 ♂ ; Ambodivoangy ; Dec. 1949 ; I. Vadon leg.; P. Vanschuytbroeck det. 1950; Sciopus ♂ tropicalis Parent; Coll. Mus. Congo; MRCA • 1 ♂ ; Andasibe , Analamazaotra Forest ; 18°46ʹ S , 48°24ʹ E ; 950 m a.s.l. ; 31 Oct.–4 Nov. 2007 ; A. Freidberg leg.; SMNHTAU • 1 ♂ ; Ambohitra , Joffreville ; 800 m a.s.l. ; 9–12 Apr. 1991 ; A. Freidberg and Fini Kaplan leg.; SMNHTAU • 1 ♂ (in glycerol); Fianarantsoa Province , Ranomafana Nat. Park ; 19 Jan. l952 ; forest; A. Pauly leg.; ex coll. RBINS . Redescription Male MEASUREMENTS. Body length 6.8 mm ; antenna length 5.6 mm ; wing length 6.8 mm ; wing width 1.9 mm . HEAD ( Fig. 3A ). Frons greenish blue-black, whitish pollinose; very short anterior vertical and long and strong postvertical bristles; upper postocular setae black, short; lateral postocular setae white, uniserial; ventral postcranium covered with irregular white long hairs; face blue-black, white pollinose, broad, 1.3 times as high as wide under antennae, 2.4 times as high as wide at clypeus, at clypeus 2.7 times as wide as postpedicel; clypeus large, slightly bulging, as high as wide; antenna ( Fig. 3B ) with scape and pedicel dirty yellow, postpedicel orange on basal half and black distally, 5 times as long as height of head; scape swollen; pedicel with short bristles, with one long dorsal and one long ventral bristles; postpedicel elongate-conoid, nearly two times as long as high, with short hairs; arista-like stylus black, apical, separated from postpedicel, with fused segments, microscopically haired; length (mm) of scape, pedicel, postpedicel, stylus, 0.09/0.1/0.21/5.33; proboscis and palpus orange, with white and black hairs; palpus with 2 black bristles. THORAX. Mesonotum and scutellum metallic bluish green, weakly pollinose; pleura bronze-green, whitish grey pollinose; metepimeron yellow; 2 long dorsocentral bristles with strongest posterior pair, and 2 hairlike setae anteriorly; acrostichals long, biserial, gradually decreasing in size anteriorly, with posterior setae as long as penultimate dorsocentrals; scutellum with 2 strong bristles and 2 minute hairs laterally. LEGS. Mostly yellow; coxae yellow, but mid coxa blackish brown from outside; fore and mid tarsi brownish to brown; hind tibia yellow to brownish; hind tarsus brown to black; fore and mid coxae with long white hairs and 2–4 black subapical bristles; hind coxa with 1 black seta at middle; femora without strong or long setae; fore femur with few ventral light hairs at base, with anterior and posterior incomplete rows of elongate setulae; mid femur with posteroventral row of elongate setulae on distal ⅔; hind femur with few long white hairs at base; fore tibia with one short posterodorsal seta at base; fore basitarsus long, without setae; all segments with ventral rows of erect and semi-erect hairs, slightly longer than width of segments, with elongate setulae on distal apex dorsally ( Fig. 3C ); mid tibia with 2 anterodorsals and 3 posterodorsals, with one short ventral seta; mid and hind tarsi simple, cylindrical; hind tibia with several short dorsal and ventral setae; femur, tibia and tarsomere (from first to fifth) length ratio (mm): fore leg: 1.83/2.19/4.92/0.99/0.71/0.37/0.21, mid leg: 1.85/3.15/3.16/0.8/0.48/0.28/0.14, hind leg: 2.65/4.23/2.15 /1.02/0.63/0.22/0.16. WING ( Fig. 3D ). Widened distally, almost hyaline, veins brown; R 4+5 gently curved to M 1 in apical third; M 1+2 straight in basal half, slightly convex anteriad in middle part; M 1 with strong elbow, forming right angle with M 1+2 ; ratio of parts of costa between R 2+3 and R 4+5 to those between R 4+5 and M 1 , 4/1; crossvein dm-m straight; ratio of crossvein dm-m to apical part of M 1+2 (fork-handle) to apical part of M 4 , 0.7/1.39/0.46; anal vein and lobe reduced; anal angle absent; lower calypter yellow, with brown cilia; halter yellow with brown knob. ABDOMEN. Thin, mostly shining blue-green, with short black hairs and long marginal setae; segment 1 yellow, with white and dark hairs; segments 2–4 each with yellow lateral spot; pregenital segments combined two times as long as mesonotum; hypopygium ( Fig. 3E ) brown basally, yellow distally, with yellow appendages; cercus with broad neck covered with hairs dorsally, 2 long arms of about equal length and one long seta between arms, half as long as arms; dorsal arm of cercus filiform, with about 8 long light setae dorsally, with one very long curved subapical seta; ventral arm of cercus with several simple and flattened setae at apex, with distal process bearing large leaf-like seta; surstylus broad, projected, bilobed; dorsal arm of surstylus deeply cleft, with 2–3 apical and subapical setae on each lobe; ventral arm of surstylus small, finger-like, with 3 small setae apically; epandrial lobe small, triangular, with 1 basal and 2 apical setae. Fig. 3. Amblypsilopus flavus ( Vanschuytbroeck, 1962 ) , ♂,Andasibe,Analamazaotra Forest (SMNHTAU). A . Head. B . Antenna. C . Fore tarsomeres 2–5. D . Wing. E . Hypopygium, left lateral view, reflected light. Distribution Madagascar (Alaotra-Mangoro, Diana, Vatovavy-Fitovinany).