Eleven new species of Amblypsilopus Bigot (Diptera: Dolichopodidae: Sciapodinae) and a key to the species of Madagascar and adjacent islands Author Grichanov, Igor Ya. text European Journal of Taxonomy 2021 2021-06-18 755 1 47 87 http://dx.doi.org/10.5852/ejt.2021.755.1399 journal article 6004 10.5852/ejt.2021.755.1399 d7d8ccff-7a58-4fc8-9eec-c2588b457d56 2118-9773 4983303 4C7C9B68-4026-45F9-98C4-430FE018119F Amblypsilopus romani sp. nov. urn:lsid:zoobank.org:act: 3E431D86-C254-44BB-B5E5-E25745BCE437 Fig. 6 Diagnosis Amblypsilopus romani sp. nov. keys to A. stuckenbergi , differing from this and other close species in the fore basitarsus distinctly shorter than fore tibia; cercus as long as epandrium, at base swollen and bearing long setae. The fore basitarsus is as long as or longer than the fore femur and tibia combined; and the cercus is distinctly elongate and setose in A. stuckenbergi and other close species of the group. Etymology The species name is dedicated to Roman Grichanov, one of my grandsons ( St. Petersburg , Russia ). Material examined Holotype MADAGASCAR • ♂ ; Andasibe , Analamazaotra Forest ; 18°46ʹ S , 48°24ʹ E ; 950 m a.s.l. ; 31 Oct.–4 Nov. 2007 ; A. Freidberg leg.; SMNHTAU . Paratypes MADAGASCAR • 2 ♂♂ ; same collection data as for holotype; SMNHTAU . Description Male ( Fig. 6A ) MEASUREMENTS. Body length 4.1 mm ; antenna length 2.5 mm ; wing length 3.8 mm ; wing width 1.1 mm . HEAD ( Fig. 6B ). Frons black (lateral view), white pollinose; strong short anterior vertical and long postvertical bristles; upper postocular setae black, short; lateral postocular setae white, uniserial; ventral postcranium covered with irregular white hairs; face black, densely white pollinose, broad, 1.5 times as high as wide under antennae, 2.7 times as high as wide at clypeus, at clypeus nearly 4 times as wide as postpedicel; clypeus small, slightly bulging; antenna ( Fig. 6C ) yellow, 2.8 times as long as height of head; pedicel with short bristles, with one long dorsal bristle; postpedicel conoid, 1.5 times as long as high, with short hairs; arista-like stylus black, brown at base, apical, microscopically haired; length (mm) of scape, pedicel, postpedicel, stylus, 0.06/0.06/0.12/2.23; proboscis yellow; palpus small, orange-yellow with white hairs and 2 black setae. Fig. 6. Amblypsilopus romani sp. nov. A–F . Holotype, ♂ (SMNHTAU). G–H . Paratype, ♂ (SMNHTAU). A . Habitus. B . Head, frontal view. C . Head and antenna, frontolateral view. D . Fore femur. E . Fore tarsus. F . Wing. G . Hypopygium, left lateral view, reflected light. H . Distal appendages of hypopygium, transmitted light. THORAX. Mesonotum orange, with brown, shining greenish blue, medial stripe expanding posteriorly; scutellum metallic blue; pleura yellow, weakly pollinose; 6 dorsocentral bristles with strongest posterior pair and 4 hair-like setae anteriorly; acrostichals biserial, decreasing in size anteriorly, with rather strong posterior pair behind suture; scutellum with 2 strong bristles and 2 hairs laterally. LEGS. Yellow; segment 5 of all tarsi black; fore and mid coxae with dark hairs and 2–3 black subapical bristles; hind coxa with 1 black bristle at middle; fore femur with 2–3 strong dorsal bristles on basal half, 1.5–2 times as long as femur height, with few anteroventral and posteroventral strong setae at base ( Fig. 6D ); fore tibia and tarsus with complete rows of semi-erect setae except ventral side, with longer and stronger setae on distal ⅓ of tibia and basal ⅓ of basitarsus, longer than diameter of podomere ( Fig. 6E ); mid tibia with two anterodorsals and one very short posterodorsal, without conspicuous ventral setae; mid and hind tarsi simple, cylindrical; hind tibia with several short dorsal and ventral setae; femur, tibia and tarsomere (from first to fifth) length ratio (mm): fore leg: 1.33/1.49/1.21/0.61/ 0.48/0.23/0.14, mid leg: 1.3/1.94/1.38/0.49/0.33/0.18/0.1, hind leg: 1.58/2.5/0.81/0.54/0.29/0.22/0.09. WING ( Fig. 6F ). Widest at middle, almost hyaline, veins brown; R 4+5 gently curved to M 1 in apical third; M almost straight; M 1 with strong elbow, forming right angle with M 1+2 ; ratio of parts of costa between R 2+3 and R 4+5 to those between R and M 1 , 5/1; crossvein dm-m straight; ratio of crossvein dm-m to apical part of M 1+2 (fork-handle) to apical part of M 4 , 0.38/0.88/0.32; anal vein foldlike, anal lobe developed; anal angle right; lower calypter brownish yellow, with fine brown cilia; halter yellow with brown knob. 1+2 4+5 ABDOMEN. Elongate, with short black hairs and marginal setae; tergite 1 yellow; tergites 2–4 each yellow, with narrow brown band along distal margin; tergites 5–6 brown; pregenital segments combined 2.1 times as long as mesonotum; hypopygium ( Fig. 6G ) dark brown, with brown appendages; cercus brownish yellow, covered with brownish hairs, somewhat longer at rounded apex, with 3 long black dorsal setae at base, longer than cercus; cerci fused at base, shorter than epandrium; hypandrium asymmetrical, with narrow left lateral arm, arising near base of hypandrium; phallus with dorsal angle; surstylus ( Fig. 6H ) projected, bilobed, with long thin arms of equal length; dorsal arm of surstylus widened at apex, with 2 short dorsoapical setae; ventral arm of surstylus with 5–6 simple long outer setae; epandrial lobe large, leaf-like, sclerotized, with pointed apex bearing few short setae; small fingerlike epandrial lobe at base of large lobe with one very long and one short seta at apex; one short epandrial ventral seta at base of epandrial lobes. Female Unknown.