Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae Author Huber, Bernhard A. text Zootaxa 2013 3713 1 1 160 journal article 10.11646/zootaxa.3713.1.1 935605ac-a0e6-4963-8264-b4b0ccb8568b 1175-5326 284803 C5F0BC11-92C0-4B30-9DB3-200882AC8950 Smeringopina bwiti new species Figs. 12 , 504–508 , 523–524 , 532 , 603–608 , 615–620 Type . ♂ holotype from Gabon , Ngounié, near Moulandoufouala ( 1°38.1’S , 10°42.5’E ), 110 m a.s.l., forest along road, 27.viii.2011 (B.A. & S.R. Huber), in ZFMK (Ar 10279). Other material examined. GABON : Ngounié : near Moulandoufouala: same data as holotype , 6♂ 6♀ 1 juv. in ZFMK (Ar 10280); same data, 1 juv. in pure ethanol, in ZFMK (Gab 187). Moyen-Ogooué : S Lambaréné near Tchad ( 0°58.1’S , 10°22.7’E ), 165 m a.s.l., forest, 27.viii.2011 (B.A. & S.R. Huber), 5♂ 1♀ in ZFMK (Ar 10281). Etymology. Named derived from bwiti , a belief system that incorporates animism, ancestor worship, and Christianity, practiced by the Babongo and Mitsogo peoples of Gabon ; noun in apposition. Diagnosis. Easily distinguished from similar congeners (large species with long abdomen, cone-shaped modified hairs on male chelicerae, simple unbranched procursus) by distinctive dorsal process on procursus ( Figs. 604 , 615–617 ), membranous ‘wings’ along second half of embolus ( Fig. 607 ), and by transversal light element ventrally on abdomen ( Fig. 523 ); also by modified male clypeus (similar to S. djidji but modified hairs on distinct humps); by shapes of male cheliceral apophyses ( Fig. 605 ; similar S. simplex but distal apophyses directed more forward), and anterior epigynal plate straight in lateral view ( Fig. 524 ; similar S. etome ). Male ( holotype ). Total body length 6.5, carapace width 1.8. Leg 1: 76.5 (18.0 + 0.8 + 17.5 + 36.7 + 3.5), tibia 2: 11.7, tibia 3: 7.7, tibia 4: 10.0; tibia 1 L/d: 104. Distance PME-PME 210 µm, diameter PME 160 µm, distance PME-ALE 70 µm, distance AME-AME 35 µm, diameter AME 160 µm. Carapace ochre with brown mark posteriorly and brown lateral margins; ocular area posteriorly brown, clypeus lower half brown, sternum frontally dark brown, posteriorly ochre-orange; legs ochre-yellow, slightly darker rings subdistally on femora and tibiae and in patella area, tips of femora and tibiae whitish; abdomen ochre-gray with distinct black pattern dorsally, laterally, and ventrally. Habitus as in Figs. 504–505 , ocular area slightly elevated, secondary eyes with distinct ‘pseudolenses’; clypeus with pair of distinct humps, each with two small modified (cone-shaped) hairs; deep thoracic pit and pair of shallow furrows diverging behind pit. Chelicerae as in Figs. 605 and 620 , with lateral proximal apophyses and distal apophyses; distal apophyses and frontal cheliceral face provided with several modified (coneshaped) hairs ( Fig. 619 ). Palps as in Figs. 506–508 ; coxa unmodified; trochanter with retrolatero-ventral apophysis; femur proximally with ventral sclerotized ridge but without pocket, with small retrolateral apophysis, without prolateral modification; prolateral femur-patella joint shifted toward ventrally (though not extremely); tarsus with some stronger hairs dorsally; procursus without (or with extremely indistinct) hinge between proximal and distal part, with distinctive dorsal process ( Figs. 603–604 , 617 ); bulb with widened but weakly sclerotized proximal part of embolus and membranous ‘wings’ along second half of embolus ( Fig. 607 ). Legs without spines and curved hairs, with few vertical hairs (many hairs missing), retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium present on all tibiae; pseudosegments barely visible. Gonopore apparently with two epiandrous spigots (not confirmed by SEM). Variation. Clypeus humps variably distinct, with 1–3 modified hairs each; dorsal process on procursus slightly variable in shape ( Figs. 615–617 ), also among specimens of same locality; sternum variably dark (entirely ochreorange to entirely brown or as in type ). Tibia 1 in 11 other males: 15.7–20.9 (mean 18.0). Female. In general similar to male; clypeus unmodified, variably dark. Tibia 1 in 7 females : 13.5–15.6 (mean 14.2). Epigynum consisting of trapezoidal anterior plate straight in lateral view and large posterior plate ( Figs. 523–524 , 606 ); internal genitalia as in Figs. 532 , 608 , 618 . Distribution. Known from two localities in western Gabon ( Fig. 468 ).