New species of spongillafly genus Sisyra Burmeister, 1839 (Neuroptera: Sisyridae) from China and Laos Author Yang, Ying Author Zheng, Yuchen Author Liu, Xingyue text Zoological Systematics 2024 49 2 129 142 http://zoobank.org/1a0dbd8f-1af9-4fde-b154-e784dc1da24f journal article 10.11865/zs.2024211 2095-6827 12172243 1A0DBD8F-1AF9-4FDE-B154-E784DC1DA24F Sisyra stenoloma sp. nov. ( Figs 6–7 ) Diagnosis. Scape, pedicel and basal two flagellomeres pale brown, following 24 flagellomeres dark brown, then remaining distal ones pale. Forewing costal space weakly broadened basally. Male sternum 9 well developed, strongly protruded postroventrally, shorter than gonocoxites 9, nearly four times as long as wide in lateral view, distally narrowed. Gonocoxites 9 long and protruded postroventrally; distal half narrowed, and twisted inward; inner margin medially with three conical tubercles, and each with a long stiff bristle. Description. Male. Body length 2.4 mm ; forewing length 3.3 mm and width 1.2 mm ; hindwing length 2.9 mm and width 1.1 mm . Body yellowish brown, but probably due to poor preservation. Head without distinct marking, covered with many long setae; compound eyes prominent. Scape pale brown, swollen medially, nearly twice as long as wide, covered with long, stout setae; pedicel pale brown, slightly longer than flagellum; flagellum with 33 segments, basal two flagellomeres pale brown, same color as scape and pedicel, following 24 flagellomeres dark brown, then remaining distal ones pale; each segment covered with many setae; apical flagellomere slightly narrow distally. Maxillary and labial palpi brown; distal segment of maxillary palpus triangular in lateral view. Thorax covered with many long setae. Legs slender; tarsomere 1 as long as total length of tarsomeres 2–4; pretarsal claws simple. Wings hyaline. Forewing ovoid, slightly curved along costal margin, distally rounded, costal space weakly broadened basally; trichosors present along distal margin; pterostigma indistinct, brownish, with crossveins; costal space with nine crossveins proximad pterostigma; two scp–ra corssveins present basally and distally; RA with three pectinate and simple branches distally; RP with three main branches; three ra–rp crossveins; MA dichotomously branched distally; one rp–ma crossvein present; MP branched into MP1 and MP2, both branches bifurcated; one ma–mp crossvein present; CuA with five pectinate and simple branches; three mp–cua crossveins; CuP, A1 and A3 simple; A2 distally bifurcated; one cua–cup, one cup–a1, one a1–a2 present. Hindwing ovoid, distally rounded; trichosors present along distal margin; pterostigma indistinct, brownish, with crossveins; costal space with six crossveins proximad pterostigma; two scp–ra corssveins present basally and distally; RP with three main branches; one ra–rp crossvein; MA dichotomously branched distally; MP branched into MP1 and MP2, both branches bifurcated; one ma–mp crossvein present, interrupted centrally by a rounded hyaline spot; CuA with six pectinate and simple branches; one mp–cua crossvein; CuP, A1 and A2 simple; one cua–cup, one cup–a1, one a1–a2 present. Figure 6. Sisyra stenoloma sp. nov. , holotype, male, China (Guangxi). A. Habitus, lateral view. B–C. Head, ventral view (left) and dorsal view (right). D–E. Fore- and hindwing. Scale bars = 1.0 mm. Figure 7. Sisyra stenoloma sp. nov. , holotype, male genitalia. A–B. lateral view. C–D. dorsal view; E–F. caudal view. Scale bars = 0.1 mm. Abdomen brown; pregenital segment with small tergum and larger sternum, covered with many long setae. Tergum 9 reduced. Sternum 9 well developed, strongly protruded postroventrally, shorter than gonocoxites 9, nearly four times as long as wide in lateral view, distally narrowed, covered with long setae. Gonocoxites 9 long and strongly protruded posteroventrad; broad basally; distal half narrowed, and twisted inward; inner margin medially with three conical tubercles, and each with a long stiff bristle. Gonocoxites 11 arched and slender, without any projection; gonostyli 11 paired, subtriangular in dorsal view, medially concaved in caudal view. Ectoprocts covered with many long setae, lateroventrally rounded; callus cerci with four trichobothria. Hypandrium internum not found. Female. Unknown. Type material. Holotype ♂ , China : Guangxi , Chongzuo , Longzhou , Nonggang , 22°28′N , 106°57′E , 20 May 1984 , Fasheng Li leg. ( CAU ). Distribution. China ( Guangxi ). Etymology. The specific epithet “ stenoloma ” refers to the strongly narrowed costal space of the forewing in the new species. Remarks. This new species has similar narrowed forewing costal space with S. yunana , but these two species can be distinguished by the antennal marking patterns and the male gonocoxites 9. In S. stenoloma sp. nov. , the scape, pedicel and basal two flagellomeres are paler than the following 24 flagellomeres, and the male gonocoxites 9 have three conical tubercles on its inner margin medially. However, in S. yunana , the basal antennomeres are in same color, and the male gonocoxites 9 bear a rounded tubercle at middle.