Phylogenetic systematics of the genera Plochionocerus Dejean and Agrodes Nordmann (Coleoptera: Staphylinidae: Xantholinini) Author Asiain, Julieta Author Márquez, Juan Author Morrone, Juan J. text Zootaxa 2007 1584 1 53 journal article 10.5281/zenodo.178410 ba2b7965-13f3-401f-9c5a-1705938112ea 1175-5326 178410 Key to the species of Plochionocerus 1. Abdominal tergites red or metallic golden; abdominal sternites metallic blue-greenish......................... 2 1’. Abdominal tergites and sternites metallic blue and/or green................................................................... 3 2(1). Head rectangular ( Fig. 2 ), dorsally and ventrally flat in lateral view; antennomeres 4–8 almost trans- verse, 9 and 10 subquadrate ( Fig. 30 ); aedeagus pear-shaped ( Fig. 92 ) ................................. P. splendens 2’. Head rounded (as in Fig. 19 ), dorsally and ventrally convex in lateral view; antennomeres 4–10 trans- verse ( Fig. 26 ); aedeagus ovate ( Fig. 82 ) ..................................................................................... P. igneus 3(1’). Head elongate, narrowed caudally; pronotal hypomeron with fine, scarce setae on anterior third; anterior margin of labrum with pair of pointed and long central teeth ( Fig. 34 ); aedeagus elongate ( Fig. 85 ) .................................................................................................................................................. P. marquezi 3’. Head not elongate, not narrowed caudally ( Figs. 1, 2 ); pronotal hypomeron lacking setae ( Fig. 63 ); anterior margin of labrum with pair of lateral teeth, variable in length, and a pair of central teeth, longer than the lateral ( Figs. 35, 36 ); aedeagus not elongate ( Figs. 77–84, 86–93 ) ........................................... 4 4(3’). Body slender; apical labial palpomere elongate, with truncate apex ( Fig. 44 ) .......................... P. gracilis 4’. Body robust; apical labial palpomere strongly widened toward obliquely truncate apex ( Fig. 43, 46, 47 ) .................................................................................................................................................................. 5 5(4’). Head rounded or ovate ( Figs. 1 , 19 ) ........................................................................................................ 6 5’. Head rectangular or subquadrate (as Fig. 2 ) .......................................................................................... 13 6(5). Dorsal surface of head slightly to strongly convex ( Figs. 11, 14 , 19 ) ..................................................... 7 6’. Dorsal surface of head flat ( Figs. 12, 13 )............................................................................................... 11 7(6). Ventral surface of head convex ( Fig. 12 )................................................................................................. 8 7’. Ventral surface of head flat ( Figs. 11, 13, 14 ) ........................................................................................ 10 8(7). Apical antennomere in males shorter than 9+10 combined (0.75–0.88 times; as in Figs. 27, 28 ); head wider than pronotum (1.31 times) ......................................................................... P. impressipennis (part) 8’. Apical antennomere in males as long as 9+10 combined (0.90–1.07 times; Figs. 25, 26 ); head as wide as or slightly wider than pronotum (1.05–1.23 times) ............................................................................. 9 9(8’). Head slightly oblong (length/width: 1.13–1.31) and slightly wider than pronotum (1.13–1.35 times); apical antennomere in males moderately longer than wide (1.36–1.54 times); ventral surface of head with dense, umbilicate punctures (20–29 on each half of head; Fig. 50 ); aedeagus ovate, parameres moderately long (0.35–0.47 times length of median lobe), apical area of median lobe 0.28–0.41 times its length and internal sac moderately visible ( Fig. 91 ) .......................................... P. simplicicollis (part) 9’. Head oblong (length/width: 1.22–1.54) and almost as wide as pronotum (0.86–1.11 times); apical antennomere in males conspicuously longer than wide (1.40–1.74 times); ventral surface of head with less dense, umbilicate punctures (10–19 on each half of head; similar to Fig. 52 ); aedeagus pear-shaped, parameres short (0.24–0.28 times length of median lobe), apical area of median lobe 0.29–0.35 times its length, internal sac clearly visible ( Fig. 79 ) ...................................................................... P. fulgens (part) 10(7’). Mandibular channel present ( Fig. 63 ); ventral surface of head with very dense, umbilicate punctures (>30 on each half of head; similar to Fig. 53 ); pronotum slightly longer than wide (1.30–1.41 times); parameres of aedeagus long (0.58–0.61 times length of median lobe; Fig. 78 ) ..................... P. discedens 10’. Mandibular channel absent or poorly developed; ventral surface of head with less dense, umbilicate punctures (10–19 on each half of head; similar to Fig. 52 ); pronotum ca. 1.5 times longer than wide (1.48–1.49 times); parameres of aedeagus moderately long (0.37–0.38 times length of median lobe; Fig. 77 ) .................................................................................................................................................. P. ashei 11(6’). Ventral surface of head moderately convex, without expanded, umbilicate punctures ( Fig. 52 ); apical antennomere in males shorter than antennomeres 9+10 combined (0.83 times their length; similar to Figs. 27, 28 ); head wider than pronotum (1.34 times); aedeagus ovate ( Fig. 80 ) ..................... P. hermani 11’. Ventral surface of head flat, with expanded, umbilicate punctures ( Figs. 50, 51, 53 ); apical antennomere in males almost as long as antennomeres 9+10 combined (0.88–1.07 times; Figs. 25, 26 ); head as wide as pronotum (1.03–1.07 times); aedeagus pear-shaped ( Figs. 87 , 93 )................................................... 12 12(11’). Mandibular channel present ( Fig. 63 ); antennomeres 4–10 moderately transverse (similar to Figs. 24, 26 ); prosternum slightly transverse (length/width: 0.72–0.80); parameres of aedeagus moderately long (0.35 times length of median lobe), internal sac clearly visible ( Fig. 87 ) .......................... P. newtonorum 12’. Mandibular channel absent; prosternum transverse (length/width: 0.61); antennomeres 4–10 strongly transverse ( Fig. 31 ); parameres of aedeagus short (0.29 times length of median lobe), internal sac moderately visible ( Fig. 93 ) ...................................................................................................... P. transversalis 13(5’). Lateroposterior half of pronotum lacking obliquely depressed areas; pronotum almost as long as elytra (1.05 times length of elytra) ................................................................................................... P. pronotalis 13’. Lateroposterior half of pronotum with two obliquely depressed areas ( Figs. 56, 57 ); pronotum shorter than elytra (<0.89 times length of elytra).............................................................................................. 14 14(13’). Apical antennomere in males shorter than antennomeres 9+10 combined (0.83–0.93 times; Figs. 27, 28 ); pronotum longer than wide (1.59–1.66 times)................................................................................ 15 14’. Apical antennomere in males as long as or longer that antennomeres 9+10 combined (> 0.96 times; Figs. 24–26 ); pronotum slightly oblong or ca. 1.5 times longer than wide (1.34–1.52 times).............. 17 15(14). Head slightly oblong (length/width: 1.19–1.25); ventral surface of head with very dense, umbilicate punctures (>30 on each half of head; Fig. 53 ); lateroposterior half of pronotum with two shallow, obliquely depressed areas ...................................................................................................... P . puncticeps 15’. Head oblong (length/width: 1.34–1.40); ventral surface of head with less dense, umbilicate punctures (10–19 on each half of head; similar to Fig. 52 ); lateroposterior half of pronotum with two clearly visible, obliquely depressed areas ( Fig. 56 )................................................................................................. 16 16(15’). Head slightly wider than pronotum (1.15–1.29 times); aedeagus with basal half curved (in lateral view), slightly longer, with internal sac moderately visible ( Fig. 84 ) .................................... P. janthinus 16’. Head wider than pronotum (1.31–1.48 times); aedeagus with basal half straight (in lateral view), slightly shorter, with internal sac conspicuously visible ( Fig. 83 ) ........................ P. impressipennis (part) 17(14’). Apical antennomere in males longer than antennomeres 9+10 combined (1.21 times; Fig. 29 ); first antennomere almost twice as long as 2+3 combined (1.88 times; Fig. 29 ) ........................... P. reticularis 17’. Apical antennomere in males almost as long as antennomeres 9+10 combined (0.96–1.06 times; Figs. 27, 30, 31 ); first antennomere 1.63–1.68 times the length of antennomeres 2+3 combined................. 18 18(17’). Head oblong (length/width: 1.26–1.50); apical antennomere in males conspicuously longer than wide (1.52–1.80 times); aedeagus pear-shaped, with apical area of median lobe 0.26–0.31 times its total length ( Figs. 79, 81 ) ............................................................................................................................... 19 18’. Head slightly oblong (length/width: 1.22–1.24); apical antennomere in males moderately longer than wide (1.42–1.45 times); aedeagus ovate, with apex of median lobe 0.39–0.41 times its total length ( Figs. 86 , 91 ) .......................................................................................................................................... 20 19(18). Antennomere 9 in males shorter than antennomere 10 (0.85 times; Fig. 25 ); ventral surface of head with dense, umbilicate punctures (20–29 on each half of head; similar to Fig. 51 ); pronotum ca . 1.5 times longer than wide (length/width: 1.47–1.58); head wider than pronotum (1.05–1.16 times); aedeagus with parameres moderately long (0.30–0.32 times length of median lobe) and internal sac moderately visible; head dorsally and ventrally flat ( Fig. 81 ) ......................................................... P. humeralis 19’. Antennomere 9 in males almost as long as antennomere 10 (0.95 times); ventral surface of head with less dense, umbilicate punctures (10–19 on each half of head; similar to Fig. 52 ); pronotum slightly oblong (length/width: 1.16–1.40); head almost as wide as pronotum (0.86–1.05 times); aedeagus with parameres short (0.24–0.28 times length of median lobe) and internal sac conspicuously visible; head dorsally and ventrally convex ( Fig. 79 ) ............................................................................ P. fulgens (part) 20(18’). Ventral surface of head with moderately dense, umbilicate punctures (10–19 on each half of head; similar to Fig. 52 ); pronotum slightly oblong (length/width: 1.34); mandibular channel absent; second antennomere 3/4 the length of third antennomere (0.75 times their length) ........................... P. modestus 20’. Ventral surface of head with dense, umbilicate punctures (20–29 on each half of head; Fig. 50 ); pronotum ca . 1.5 times longer than wide (length/width: 1.47); mandibular channel present ( Fig. 63 ); second antennomere ca . 1/2 the length of third antennomere (0.52–0.68 times its length)... P. simplicicollis (part)