Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae) Author Pall-Gergely, Barna Author Budha, Prem B. Author Naggs, Fred Author Backeljau, Thierry Author Asami, Takahiro text ZooKeys 2015 529 1 70 http://dx.doi.org/10.3897/zookeys.529.6139 journal article http://dx.doi.org/10.3897/zookeys.529.6139 1313-2970-529-1 AD4323B4913C447A88A7CE05EC8862A3 Taxon classification Animalia Pulmonata Plectopylidae Endothyrella oglei (Godwin-Austen, 1879) Figures 5A, 6A Endothyrella oglei 1879a Helix (Plectopylis) Oglei Godwin-Austen: Journal of the Asiatic Society of Bengal, 48 (2): 3, Plate 1, figs 2, 2 a-c . ["Near Sadiya, Assam"]. Endothyrella oglei 1887 Helix (Plectopylis) oglei , - Tryon: Manual of Conchology ... , 2 (3): 159, Plate 36, figs 29-31. Endothyrella oglei 1898 Plectopylis oglei , - Gude: Science Gossip, 4: 263, figs 68 a-h . Endothyrella oglei 1899c Plectopylis (Chersaecia) oglei , - Gude: Science Gossip, 6: 148. Endothyrella oglei 1899d Plectopylis (Chersaecia) oglei , - Gude: Science Gossip, 6: 175, 176. Endothyrella oglei 1914b Plectopylis (Chersaecia) oglei , - Gude: The Fauna of British India ... : 73, 92-93, figs 39 a-h . Types. Sadia, E. Assam, leg. Ogle, NHMUK 1903.7.1.740. (4 syntypes, Figure 5A, 6A). Figure 5. Shells of Endothyrella species. A Endothyrella oglei (Godwin-Austen, 1879), NHMUK 1903.7.1.740. (syntype) B Endothyrella serica (Godwin-Austen, 1875), NHMUK 1903.7.1.741 (syntype of serica ) C Endothyrella serica , NHMUK 1903.7.1.744 (syntype of serica ) D Endothyrella serica , NHMUK 1903.7.1.742. (syntype of munipurensis ). All photos by Harold Taylor (NHMUK). Scale represent 5 mm. Figure 6. Photos ( A-C ) and SEM images ( D-F ) of Endothyrella protoconchs. A Endothyrella oglei (Godwin-Austen, 1879), same data as in Fig. 5 B Endothyrella serica (syntype of munipurensis , same data as in Fig. 5.) C Endothyrella babbagei (Gude, 1915), same data as in Fig. 3 D Endothyrella plectostoma (Benson, 1836), MNHN 2012-27053 E Endothyrella nepalica Budha & Pall-Gergely sp. n., paratype from the Siddha Cave F Endothyrella inexpectata Pall-Gergely , sp. n., (holotype). A-C Harold Taylor D-F B. Pall-Gergely . Figure 7. Distribution of Endothyrella species. Empty circle: Endothyrella inexpectata sp. n.; Filled triangle, top up: type locality of Endothyrella babbagei . Letters indicate localities of Endothyrella plectostoma (Benson, 1836). Abbreviations: A Arakan HillsB Bassein (= Pathein)C Silchar (Cachar)Ch Chittagong (Ghoramara)D DarjeelingDu Dunsiri valleyG Garo HillsK Khasi HillsM ManipurN Naga HillsP Pegu (= Bago)S SylhetSw ShweguY Ywathit. Figure 8. SEM images of Endothyrella shells. A Endothyrella nepalica Budha & Pall-Gergely , sp. n., 4th, 5th whorl, for locality see Fig. 6. B Endothyrella nepalica Budha & Pall-Gergely , sp. n. body whorl C Endothyrella nepalica Budha & Pall-Gergely , sp. n. body whorl D Endothyrella inexpectata Pall-Gergely , sp. n., body whorl (holotype). All images by B. Pall-Gergely . Figure 9. Parietal (A, C, E, G, I, K) and palatal (B, D, F, H, J, L) lamellation of Endothyrella spp. A-B Endothyrella inexpectata sp. n. (holotype) C-D Endothyrella nepalica sp. n., for locality, see Fig. 6. E-F Endothyrella nepalica sp. n., shell from Mahadevsthan G-H Endothyrella dolakhaensis sp. n., shell from the type locality I-J Endothyrella angulata sp. n., shell from the type locality K-L Endothyrella robustistriata sp. n., NHMUK 1903.7.1.767 (two different specimens). Outer view: B, L; inner view: D, F, H, J. Diagrammatic. Diagnosis. Shell middle sized, dextral, yellowish-reddish striped with moderately wide umbilicus and somewhat domed dorsal surface; callus strong, palatal plicae divided at their middle and the posterior fragments are connected by a ridge; parietal wall with a single curved lamella with posteriorly elongated upper and lower ends. Probably at least the upper elongation is homologous with the posterior denticle of other Endothyrella species. Measurements (in mm): D: 16.8-16.9, H: 7.7-8.1 (n = 2, type series). Differential diagnosis. Endothyrella babbagei and Endothyrella inexpectata sp. n. differ from the Endothyrella oglei by the flat dorsal surface of the shell and the presence of hairs arranged in three rows on the body whorl. See also under Endothyrella serica and Table 5. Remarks. The information published by Gude (1914b) (major diameter 27, minor diameter 25 mm) is wrong; it probably refers to " Chersaecia " andersoni . Distribution. The species is known from the type locality only (Figure 10). Figure 10. Distribution of Endothyrella species in Northeastern India. Abbreviations: A type locality of Endothyrella aborensis (Gude, 1915); Abor Abor Hills (type locality of Endothyrella williamsoni (Gude, 1915)BE type locality of Endothyrella bedfordi (Gude, 1915)BR Type locality of Endothyrella brahma (Godwin-Austen, 1879); Miri Miri Hills (type locality of Endothyrella miriensis (Gude, 1915)OA Type locality of Endothyrella oakesi (Gude, 1915)OG Type locality of Endothyrella oglei (Godwin-Austen, 1879).