<document id="705C18BF0B53F791A7819791AC7F55C3" ID-CLB-Dataset="30595" ID-DOI="10.11646/zootaxa.4388.4.1" ID-GBIF-Dataset="a884362d-a9f8-4cee-8cd4-dcf59a73ede2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1189072" ID-ZooBank="7A57C29B-6B16-445E-A8D7-750C736E6B42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520318467094" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2018" docId="5C282C04FFCFFFEDFF2D57C81AD1FB87" docLanguage="en" docName="zootaxa.4388.4.1.pdf" docOrigin="Zootaxa 4388 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthidium (Canthidium) variolosum Howden &amp; Young 1981" docType="treatment" docVersion="10" lastPageNumber="465" masterDocId="A011547CFFC1FFE3FFBA541B1E1AFFFB" masterDocTitle="On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species" masterLastPageNumber="486" masterPageNumber="451" pageNumber="465" updateTime="1698528354521" updateUser="ExternalLinkService">
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<mods:title id="2931AF63E401617B20E4B168E26AA8EF">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<taxonomicName id="1381E691FFCFFFEDFF2D57C81D6AFC16" ID-CoL="68XF8" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,880,979,1005]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="variolosum" subGenus="Canthidium">
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<emphasis id="E6F54100FFCFFFEDFF2D57C81C57FC17" bold="true" box="[151,589,979,1004]" italics="true" pageId="14" pageNumber="465">Canthidium (Canthidium) variolosum</emphasis>
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<emphasis id="E6F54100FFCFFFEDFF7D57E31FCEFBEB" box="[199,468,1016,1040]" italics="true" pageId="14" pageNumber="465">Canthidium variolosum</emphasis>
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: 88
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[here transferred to the subgenus 
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]. 
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locality:
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<collectingCountry id="AC96DD82FFCFFFEDFF4750061F40FBCF" box="[253,346,1053,1076]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
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: 
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<collectingCountry id="AC96DD82FFCFFFEDFEDF50061FD8FBCF" box="[357,450,1053,1076]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
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: San Carlos: 
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, 
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west of Hato del Volcán (“ 
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<collectingCountry id="AC96DD82FFCFFFEDFB8750071A86FBCF" box="[1085,1180,1052,1076]" name="Panama" pageId="14" pageNumber="465">Panamá</collectingCountry>
</collectingRegion>
, Las Lagunas, 
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W Hato del Volcán”). 
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<typeStatus id="0B3A23B0FFCFFFEDFECE50241FB5FBA3" box="[372,431,1087,1112]" pageId="14" pageNumber="465">Type</typeStatus>
material:
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<typeStatus id="0B3A23B0FFCFFFEDFD9550241C8DFBA3" box="[559,663,1087,1112]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
male (CMNC—ex Howden collection); see full list of 
<typeStatus id="0B3A23B0FFCFFFEDFAA8505B1B67FBA3" box="[1298,1405,1088,1112]" pageId="14" pageNumber="465" type="paratype">paratypes</typeStatus>
in 
<treatmentCitation id="5520BB03FFCFFFEDFF2D507F1FF8FB87" author="" box="[151,482,1124,1149]" page="88" pageId="14" pageNumber="465">
<bibRefCitation id="B010E0E3FFCFFFEDFF2D507F1FF8FB87" box="[151,482,1124,1149]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young (1981: 88)</bibRefCitation>
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. 
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and 
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<collectingCountry id="AC96DD82FFCFFFEDFCFA507E1D83FB87" box="[832,921,1125,1148]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
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(Solís &amp; Kohlmann 2004).
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