<document ID-CLB-Dataset="209209" ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-GBIF-Dataset="bac17de6-baf9-4fee-9ded-268adbb26a61" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo" docDate="2022" docId="3E8350C76DFC51EFA098A086FA0FF2B2" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Toliaranella fisheri Lorenz & Loria & Harvey & Harms 2022, sp. nov." docType="treatment" docVersion="5" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1732733992417" updateUser="ExternalLinkService"> <mods:mods id="318B07BB6FC2D24F8E9E01A84BE910DB" xmlns:mods="http://www.loc.gov/mods/v3"> <mods:titleInfo id="716F93C7C27247D24C6BCDE8C2BE2035"> <mods:title id="AC168DA9C4D0F5C3A9A78B43A124CA38">The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots</mods:title> </mods:titleInfo> <mods:name id="744EB7F58C945B1C8065EBEED63449B1" type="personal"> <mods:role id="EB5AE65543D2B5917A6E01DAB5928C1A"> <mods:roleTerm id="BD89BA98405F51A2C629E9F903D7536F">Author</mods:roleTerm> </mods:role> <mods:namePart id="4A0A1A6F59181B2B30B861CE6EB013EA">Lorenz, Michelle</mods:namePart> <mods:affiliation id="77F21DDD395C96A01456BDEEB322DD74">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation> <mods:nameIdentifier id="46A56D9B7924944A9A4D234CDE951DAE" type="email">michellelorenz@gmx.net</mods:nameIdentifier> </mods:name> <mods:name id="FC1BE803CE900018A80C789B4982ECE5" type="personal"> <mods:role id="EFA6EA4FC3E329E76F3F400424E3E556"> <mods:roleTerm id="5436CB2DFB28C35500ED6D0AB5D83985">Author</mods:roleTerm> </mods:role> <mods:namePart id="1149EAF12F11997D73F25557A510DBA4">Loria, Stephanie F.</mods:namePart> <mods:affiliation id="ADF43CFDCECAC235276B0EC619DE0D3E">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation> </mods:name> <mods:name id="8DC6551948BC8DA569E3E7F4B3249BA2" type="personal"> <mods:role id="4365D2D1D6900E54564F2C13D26A62C9"> <mods:roleTerm id="891E7C17E2E9730608370DEB17CC6DAF">Author</mods:roleTerm> </mods:role> <mods:namePart id="83E477C0EF9CBF00608C691D5F9B305D">Harvey, Mark S.</mods:namePart> <mods:nameIdentifier id="515EDF9E3964A695751004360CD8E51A" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier> <mods:affiliation id="86035143345A77096C532117EDE80B7D">Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, WA 6106, Australia & University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation> </mods:name> <mods:name id="2C703BB262ADFA30F8A274B6931F655C" type="personal"> <mods:role id="0B736796F560375EDC6E1253811392D5"> <mods:roleTerm id="746C9EB91D0FCC84E30D719C42C29BDF">Author</mods:roleTerm> </mods:role> <mods:namePart id="A00366CEEEF75B8980D87A4B02024B2D">Harms, Danilo</mods:namePart> <mods:nameIdentifier id="AB6B7AD448D43150DDFE6B52F6D33B54" type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier> <mods:affiliation id="9C8D0DA02C40DCEE67A2869AB075DC6B">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation> </mods:name> <mods:typeOfResource id="6FE6E758A0A7E6F6230EA5D2F56685A9">text</mods:typeOfResource> <mods:relatedItem id="16F91EDF490839F1E7153FF5B9ABDBB1" type="host"> <mods:titleInfo id="7A90878B810E16614FEBF26E7500718C"> <mods:title id="0838EAA703359DE0BA195409F4E08F89">Arthropod Systematics & amp; Phylogeny</mods:title> </mods:titleInfo> <mods:part id="0EB5CB88D0111A40D143B312A06D787F"> <mods:date id="314AC4484CE026BC47E8635906B85B26">2022</mods:date> <mods:detail id="862580655603CA991D6E6BF679F55A87" type="pubDate"> <mods:number id="8844F9DB500C65D16FF7B8086C88BDBB">2022-12-14</mods:number> </mods:detail> <mods:detail id="E19BE47D186C09A337EA68C63F02646E" type="volume"> <mods:number id="365E22901505782073308A9E308412E7">80</mods:number> </mods:detail> <mods:extent id="86AAB4FF881A8F0836863A3E2D16058F" unit="page"> <mods:start id="6FFFD8AABC6453A4D36639BA47D7812C">649</mods:start> <mods:end id="9430E99F93542E64C1651137C939C247">691</mods:end> </mods:extent> </mods:part> </mods:relatedItem> <mods:location id="AFD3343A7DB32664421519BEDB93C388"> <mods:url id="7649138CED34162FC6BA3C2D0ABECFEA">http://dx.doi.org/10.3897/asp.80.e90570</mods:url> </mods:location> <mods:classification id="1DE39B80DBB5E45EF764C168FD92BB18">journal article</mods:classification> <mods:identifier id="A2CD948F1B23C3E8DB494C330A70E757" type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier> <mods:identifier id="2BB0BE0EBF5B61400A63F8688A32F5D1" type="Pensoft-Pub">1864-8312-80-649</mods:identifier> <mods:identifier id="A74823601976AA9E00B0A66417251B1A" type="ZooBank">AD0B1CC262414460B6B888E6B95F20D7</mods:identifier> <mods:identifier id="AE4FCF946A4E92FEA39BACE637F7974E" type="Pensoft-UUID">FAE6DFEF69735531A4CBA275E078320D</mods:identifier> </mods:mods> <treatment id="3E8350C76DFC51EFA098A086FA0FF2B2" ID-GBIF-Taxon="206273802" LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/D3AA212A-7763-41C7-A683-62F9FAC74FFC" httpUri="http://treatment.plazi.org/id/3E8350C76DFC51EFA098A086FA0FF2B2" lastPageNumber="649" pageId="0" pageNumber="649" scope_family="Feaellidae" scope_order="Pseudoscorpiones" scope_subFamily="Feaellinae" scope_superFamily="Feaelloidea"> <subSubSection id="E31650B4AB92BEEE1F4A8B57044B89B4" pageId="0" pageNumber="649" type="nomenclature"> <paragraph id="CBD57F818AE1090FDD125685F4809BB9" pageId="0" pageNumber="649"> <taxonomicName id="DBF6F6E03E28A00A96713374612B2002" LSID="https://zoobank.org/NomenclaturalActs/D3AA212A-7763-41C7-A683-62F9FAC74FFC" authority="Lorenz & Loria & Harvey & Harms, 2022" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella fisheri" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">Toliaranella fisheri</taxonomicName> <taxonomicNameLabel id="6A72DB4E9D858D401D56E885A798505D" pageId="0" pageNumber="649">sp. nov.</taxonomicNameLabel> </paragraph> </subSubSection> <subSubSection id="2170A18C9F653DF5E2C032E81EA27D9D" pageId="0" pageNumber="649" type="type material"> <paragraph id="8C7BCE6F602F2F628C4904A9547FBD41" pageId="0" pageNumber="649">Type material examined.</paragraph> <paragraph id="D26E88CE4CDF060490855FA31A08AF02" pageId="0" pageNumber="649"> MADAGASCAR: Toliara Province [Anosy Region]: Holotype ♂ (CAS 9071590 [BLF5316]), allotype ♀ (ZMH-A0016730), paratypes: 6 ♂, 7 ♀, (CAS 9071590 [BLF5316]), 4 ♂, 4 ♀ (ZMH-A0016731 - ZMH-A0016738), Reserve Prive Berenty, Foret de Bealoka, <geoCoordinate id="F92D5F5EC6135808AFC96689197AF791" degrees="24" direction="south" minutes="57" orientation="latitude" precision="15" seconds="25" value="-24.956945">24°57′25″S</geoCoordinate> <geoCoordinate id="1E451100071FCDECAB23BA3C6CA1D568" degrees="46" direction="east" minutes="16" orientation="longitude" precision="15" seconds="17" value="46.27139">46°16′17″E</geoCoordinate> , 3-8 February 2002, B.L. Fisher, Griswold et al. </paragraph> </subSubSection> <subSubSection id="6C9353845BEAC38CC93496CDA497C153" pageId="0" pageNumber="649" type="materials_examined"> <paragraph id="A465389BD80EEAA61375C5639FD47AE6" pageId="0" pageNumber="649">Additional material examined.</paragraph> <paragraph id="1C4FB0572CACF347E91B6E97A1AAF676" pageId="0" pageNumber="649"> <materialsCitation id="D1E6130F86EB7C6EAAD7C9E00AF7E4BC" ID-GBIF-Occurrence="3990971324" collectingDate="2002-02-03" collectingDateMax="2002-02-08" collectingDateMin="2002-02-03" collectorName="de Bealoka, B. L. Fisher, Griswold" country="MADAGASCAR" latitude="-24.956945" location="Foret de Bealoka" longLatPrecision="21" longitude="46.27139" municipality="Reserve Prive Berenty" specimenCount="1" specimenCount-male="1" stateProvince="Toliara Province"> <collectingCountry id="76ABB73F9413B4BC8D3A42DB965B19D3" name="Madagascar">MADAGASCAR</collectingCountry> : <collectingRegion id="86349385C5F506AA96801A547D8CA862" country="Madagascar" name="Toliara">Toliara Province</collectingRegion> [Anosy Region]: <specimenCount id="AC9D31F80FAA27CF44EC3C0EBD6E8B8A" type="male">1 ♂</specimenCount> , 1 nymph (CAS 9071750 [BLF5314]), <collectingMunicipality id="11C128572EB94E42BAAD13A376D01C3A">Reserve Prive Berenty</collectingMunicipality> , <location id="44E12D79FFCD8474B3CEB1FAA8ABA0FC" LSID="urn:lsid:plazi:treatment:3E8350C76DFC51EFA098A086FA0FF2B2:44E12D79FFCD8474B3CEB1FAA8ABA0FC" country="MADAGASCAR" latitude="-24.956945" longLatPrecision="21" longitude="46.27139" municipality="Reserve Prive Berenty" name="Foret de Bealoka" stateProvince="Toliara Province"> Foret <collectorName id="0AF4A44841687C6F6E87156E5B98325B">de Bealoka</collectorName> </location> , <geoCoordinate id="5D6F49D65A9EDCC8AA74BA3B63412CE8" degrees="24" direction="south" minutes="57" orientation="latitude" precision="15" seconds="25" value="-24.956945">24°57′25″S</geoCoordinate> <geoCoordinate id="45EE73BE11D12A7D87E9FC48CDD263D4" degrees="46" direction="east" minutes="16" orientation="longitude" precision="15" seconds="17" value="46.27139">46°16′17″E</geoCoordinate> , <collectingDate id="C8C7AEF54F89219F5E18230389C52D35" value="2002-02-03" valueMax="2002-02-08" valueMin="2002-02-03">3-8 February 2002</collectingDate> , <collectorName id="316DDE00F346A3833B3EC54634E8C401">B.L. Fisher</collectorName> , <collectorName id="6557BBE9B57B9A19A2F21EEA32506175">Griswold</collectorName> et al. 76 </materialsCitation> ; <materialsCitation id="37B71B46802B52543BA6BE6CFA9A7526" ID-GBIF-Occurrence="3990971329" collectingDate="2002-02-06" collectorName="Fisher, Griswold" country="Madagascar" county="Reserve Prive Berenty" latitude="-25.007778" location="Mandrare River" longLatPrecision="21" longitude="46.30611" municipality="Foret de Malaza" specimenCount="55" specimenCount-female="55" stateProvince="Toliara Province"> <specimenCount id="07672FD8DA07C352D78671D3FA66CAE3">♂</specimenCount> , <specimenCount id="51B8A3C43CDD8CBCA9E005C7968B691A" type="female">55 ♀</specimenCount> ,>100 nymphs (CAS 9071589 [BLF5426]), <collectingCounty id="BA729CDF58B743DF7C5D9A996E7A39CC">Reserve Prive Berenty</collectingCounty> , <collectingMunicipality id="221227BAAC43A8612ECAD7EF41E7091A">Foret de Malaza</collectingMunicipality> , <location id="24EDD2EA4FC58E9FEEC103E03C0FEB30" LSID="urn:lsid:plazi:treatment:3E8350C76DFC51EFA098A086FA0FF2B2:24EDD2EA4FC58E9FEEC103E03C0FEB30" country="Madagascar" county="Reserve Prive Berenty" latitude="-25.007778" longLatPrecision="21" longitude="46.30611" municipality="Foret de Malaza" name="Mandrare River" stateProvince="Toliara Province">Mandrare River</location> , <geoCoordinate id="01178E5784E5A2DD81B3B41240C225F1" degrees="25" direction="south" minutes="00" orientation="latitude" precision="15" seconds="28" value="-25.007778">25°00′28″S</geoCoordinate> <geoCoordinate id="6D1501F631F203D10E54FF8FCF531377" degrees="46" direction="east" minutes="18" orientation="longitude" precision="15" seconds="22" value="46.30611">46°18′22″E</geoCoordinate> , <collectingDate id="59E0BC04188DDBBCC07FFFF611E727DD" value="2002-02-06">6 February 2002</collectingDate> , <collectorName id="110B8E488AFADA10787943831A8BC92C">Fisher</collectorName> , <collectorName id="B9F10A727FE71486450FEB69518FEA50">Griswold</collectorName> et al </materialsCitation> .; <materialsCitation id="685B7C26525E41898460F81DED1769E4" ID-GBIF-Occurrence="3990971307" collectingDate="2002-01-31" collectorName="Fisher, Griswold" country="Madagascar" latitude="-24.75361" location="Isantoria River" longLatPrecision="21" longitude="46.15139" municipality="Foret de Mahavelo" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toliara Province"> <specimenCount id="DCDA8EBB731391C52B431AE2536C8AF4" type="male">1 ♂</specimenCount> , <specimenCount id="F52829C6671C25A6A76FD8F21EB76E0E" type="female">2 ♀</specimenCount> , 2 nymphs (CAS 9071578 [BLF5278]), <collectingMunicipality id="787F5D20C2B1A4EE129B06590E617505">Foret de Mahavelo</collectingMunicipality> , <location id="78D63FC845351B35C1313BDC99BCC997" LSID="urn:lsid:plazi:treatment:3E8350C76DFC51EFA098A086FA0FF2B2:78D63FC845351B35C1313BDC99BCC997" country="Madagascar" latitude="-24.75361" longLatPrecision="21" longitude="46.15139" municipality="Foret de Mahavelo" name="Isantoria River" stateProvince="Toliara Province">Isantoria River</location> , <geoCoordinate id="6DAD788F0542F4FE24AC1D7050FC2FEA" degrees="24" direction="south" minutes="45" orientation="latitude" precision="15" seconds="13" value="-24.75361">24°45′13″S</geoCoordinate> <geoCoordinate id="F4C114E24421CB629B0FF6F6B52C7805" degrees="46" direction="east" minutes="09" orientation="longitude" precision="15" seconds="5" value="46.15139">46°09′5″E</geoCoordinate> , <collectingDate id="FDD021040B96085DEE5E25B9A87005EC" value="2002-01-31">31 January 2002</collectingDate> , <collectorName id="A5E0B8DB62E06E7EE116C0E792C15C41">Fisher</collectorName> , <collectorName id="6B7D35E38E3D6CC632F31B9C40136201">Griswold</collectorName> et al </materialsCitation> . </paragraph> </subSubSection> <subSubSection id="75D13D0AFD1B26BAADDAF31C2C2CB741" pageId="0" pageNumber="649" type="diagnosis"> <paragraph id="BE7082227304E772633215EFD43E7E53" pageId="0" pageNumber="649">Diagnosis.</paragraph> <paragraph id="1AEFB602F80162080FB5551CB0B74C9C" pageId="0" pageNumber="649"> Most similar to <taxonomicName id="C7C95EF47B6320C76A1F05C6D6BBD364" lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi"> <emphasis id="071AAED4F5B7D322D1B59341A8E51BE7" italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis> </taxonomicName> <emphasis id="EC7F16F09B94DC3013A19421AF0018BE" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis> , but females with most slender femora of this new genus (1.71 <normalizedToken id="66031941ABD2817BC8E787D75537F3D5" originalValue="×">x</normalizedToken> longer than broad versus 1.62-1.69) and different body size. </paragraph> </subSubSection> <subSubSection id="AE66FAFBE20BB54099C68E15EE3ED373" pageId="0" pageNumber="649" type="etymology"> <paragraph id="FA0C9FAAF463C42D527D6A528DF9E512" pageId="0" pageNumber="649">Etymology.</paragraph> <paragraph id="FC22578AF34CAA8C4C25E332F9F93406" pageId="0" pageNumber="649">This species is a patronym honoring Brian Fisher who collected many Malagasy feaellids.</paragraph> </subSubSection> <subSubSection id="F4394ED9695A4BC528994DCC7FA2A23E" pageId="0" pageNumber="649" type="description"> <paragraph id="45A556D88AF005A183487B8A67FF02F6" pageId="0" pageNumber="649">Description.</paragraph> <paragraph id="8BEB6A083C3AE577E90A7F363D9B2C89" pageId="0" pageNumber="649"> The following description is based on holotype and allotype. - <emphasis id="5D68B67C2770F77BBE0183D4A2F75A0C" italics="true" pageId="0" pageNumber="649">Carapace</emphasis> (Fig. <figureCitation id="4446AFC26CFF3BF3DF3B9BE011F02282" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21A</figureCitation> ): 1.25-1.39 (♂), 1.27-1.39 (♀) times longer than broad. - <emphasis id="BAD3A6055E53C45526D4EFB7F75865CC" italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis> (Fig. <figureCitation id="A5D9A4654352F8843AC0FA45E183B1BF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21C, D</figureCitation> ): See trichobothrial pattern in description of <taxonomicName id="CB55FAFD5DAF2F14D9FA5EC3E3663F6D" lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi"> <emphasis id="DC93E4B361D80AC437989349E3AFCBFB" italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis> </taxonomicName> <emphasis id="4F94A1C077F839015CE6CF50039B93FE" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis> Chelal fixed finger with 10 teeth in the OR, 13-16 in the MR and 10-13 teeth in the IR; movable finger with 8-10 teeth in the OR, 9-12 in the MR and 8-11 teeth in the IR. Fixed finger with 6-7 equally sized and one large terminal tooth, movable finger with 7 terminal teeth equal in size. - <taxonomicName id="167C05A222B5BD866AE48045A7671DF3" genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus"> <emphasis id="0EFC44AE758F46C7C6F01F71643E3FCD" italics="true" pageId="0" pageNumber="649">Dimensions</emphasis> </taxonomicName> (mm): <emphasis id="6F381BB36C21973C4541D744754BF769" italics="true" pageId="0" pageNumber="649">Holotype</emphasis> ♂: Body length 1.56; abdomen length 1.10, width (with pleura) 1.00, width (without pleura) 0.85; carapace length 0.44, width 0.34. Pedipalp: trochanter 0.24; femur length 0.43, width 0.27; patella 0.36; chela (without pedicel) 0.39; hand length 0.09, width 0.12; movable finger 0.32. Leg I: trochanter 0.07; femur 0.16; patella 0.14; tibia; 0.13; tarsus 0.22. Leg IV: trochanter 0.15; femur 0.10; patella 0.25; tibia 0.26; tarsus 0.26. <emphasis id="9EF61F7CF88A23FA8CBD0455E0D9A97D" italics="true" pageId="0" pageNumber="649">Allotype</emphasis> ♀: Body length 2.15; abdomen length 1.49, width (with pleura) 1.35, width (without pleura) 1.13; carapace length 0.57, width 0.44. Pedipalp: trochanter 0.24; femur length 0.60, width 0.33; patella 0.45; chela (without pedicel) 0.50; hand length 0.11, width 0.15; movable finger 0.41. Leg I: trochanter 0.10; femur 0.20; patella 0.16; tibia 0.15; tarsus 0.28. Leg IV: trochanter 0.19; femur 0.13; patella 0.30; tibia 0.31; tarsus 0.35. </paragraph> <caption id="5FA615B43D59BF0ACD596FBCF2D65E3B" doi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649" start="Figure 21" startId="F21"> <paragraph id="9152644DC9788CCA3A3782C33E0A9328" pageId="0" pageNumber="649"> <emphasis id="CA25FF8BDA0A2DB363ED91B34191D3A3" bold="true" pageId="0" pageNumber="649">Figure 21.</emphasis> Drawings of <taxonomicName id="98C093A07F54796F9265C3584C6D553F" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella fisheri" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="fisheri"> <emphasis id="AD5B8942E81AEFE2EC5F9714682E9886" italics="true" pageId="0" pageNumber="649">Toliaranella fisheri</emphasis> </taxonomicName> <emphasis id="4BDA18C264120E08180EDB632F2E4F42" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis> : <emphasis id="25B271087D08F65FF5ACC7FCF8210D8A" bold="true" pageId="0" pageNumber="649">A</emphasis> carapace from dorsal; <emphasis id="ACA891CCC89CB7BBE3C5D31DB177E65D" bold="true" pageId="0" pageNumber="649">B</emphasis> leg IV; <emphasis id="EE7DFD43F72153E3239554940A5DC0A4" bold="true" pageId="0" pageNumber="649">C</emphasis> left chela from dorsal with trichobothrial pattern; <emphasis id="CDF4EC56386AF741BF1236D8F1327F7D" bold="true" pageId="0" pageNumber="649">D</emphasis> right pedipalp from dorsal. Scale bars: 0.2 mm ( <emphasis id="D43D8A61F4F297FAF3980A2487BEAAFD" bold="true" pageId="0" pageNumber="649">A</emphasis> - <emphasis id="42C7E1CFB1A4F65160AE7A86593E2D3F" bold="true" pageId="0" pageNumber="649">C</emphasis> ); 0.4 mm ( <emphasis id="D32E4DD4638EFAED4A34CE34E23E266D" bold="true" pageId="0" pageNumber="649">D</emphasis> ). </paragraph> </caption> </subSubSection> <subSubSection id="D034D9A41806BDB70EA2A9BCCD63CC99" pageId="0" pageNumber="649" type="variation"> <paragraph id="D815D1E31D17565DAB25B7D4C3904691" pageId="0" pageNumber="649">Variation.</paragraph> <paragraph id="A598F149C529F5AEC8B9FAED5114730B" pageId="0" pageNumber="649">Body length 1.54-1.72 (♂), 1.83-2.15 (♀); abdomen length 1.07-1.16 (♂), 1.30-1.54 (♀); width (with pleura) 0.97-1.05 (♂), 1.16-1.38 (♀); width (without pleura) 0.83-0.92 (♂), 1.00-1.17 (♀); 1.23-1.29 (♂), 1.27-1.40 (♀) times longer than broad; carapace length 0.44-0.50 (♂), 0.51-0.58 (♀); width 0.34-0.38 (♂), 0.39-0.44 (♀); 1.25-1.39 (♂), 1.27-1.39 (♀) times longer than broad. Pedipalp: trochanter 0.23-0.27 (♂), 0.22-0.27 (♀); femur length 0.43-0.48 (♂), 0.51-0.60 (♀); width 0.26-0.29 (♂), 0.29-0.35 (♀); 1.59-1.78 (♂), 1.56-1.84 (♀) times longer than broad; patella 0.36-0.39 (♂), 0.41-0.47 (♀); chela (without pedicel) 0.39-0.43 (♂), 0.44-0.50 (♀); hand length 0.08-0.11 (♂), 0.10-0.12 (♀); width 0.11-0.13 (♂), 0.13-0.15 (♀); 0.67-0.92 (♂), 0.67-0.86 (♀) times longer than broad; movable finger 0.31-0.35 (♂), 0.36-0.41 (♀).</paragraph> </subSubSection> <subSubSection id="B2A9FD937848AD5D569E7ED12CC097CB" pageId="0" pageNumber="649" type="genbank code"> <paragraph id="646D5E4E0483B9E627BBD8DE212406AE" pageId="0" pageNumber="649">GenBank Code.</paragraph> <paragraph id="B60D331B062F18A061D065B3F340E0A8" pageId="0" pageNumber="649">OP589962, OP589963. The species differs from all congeners by more than 3.7% in the H3 dataset.</paragraph> </subSubSection> <subSubSection id="04B3B36CD01EB229CE877B63E5714130" pageId="0" pageNumber="649" type="habitat"> <paragraph id="B46A2DDC731667D347CCD778E06C46A8" pageId="0" pageNumber="649">Habitat.</paragraph> <paragraph id="E2EDD1A462FFE0A489F96862DF7F9EAC" pageId="0" pageNumber="649">Specimens were found in sifted litter (leaf mold and rotten wood) in gallery forest and spiny forest thicket at an elevation of 35-115 m.</paragraph> </subSubSection> <subSubSection id="428C88D7BFAD091AAFB6321AF880776C" pageId="0" pageNumber="649" type="distribution"> <paragraph id="84AE8E54763FD6D47CA876153B0AF969" pageId="0" pageNumber="649">Distribution.</paragraph> <paragraph id="70A216AB475B07AF654776D3AE05B746" pageId="0" pageNumber="649">Known from three localities along the Mandrare River in the Anosy Region (formerly Toliara Province) in southern Madagascar.</paragraph> </subSubSection> </treatment> </document>