On Venezuelan pholcid spiders (Araneae, Pholcidae)
Author
Huber, Bernhard A.
33607F65-19BF-4DC9-94FD-4BB88CED455F
Zoological Research Museum Alexander Koenig, Bonn, Germany.
b.huber@leibniz-zfmk.de
Author
Villarreal, Osvaldo
679C385E-B068-4351-9D2F-97753E534C26
Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. & Museu Nacional / UFRJ, Rio de Janeiro, Brazil.
osvaldovillarreal@gmail.com
text
European Journal of Taxonomy
2020
2020-10-01
718
1
317
journal article
10.5852/ejt.2020.718.1101
4069574
F9E9A91E-488C-4DB1-9361-E788E9AC5BC1
Canaima guaquira
Huber
sp. nov.
urn:lsid:zoobank.org:act:
ABAD982F-9BF4-4CE6-BF7D-934296A0224E
Figs 54–55
,
85–88
,
96–98
, 1034
Diagnosis
Distinguished from known congeners by armature of male chelicerae (
Fig. 87
; three pairs of processes, with highly distinctive distal pair of slender curved processes), by shape of procursus (
Fig. 86
; distinctive dorso-distal sclerite), by dorsal process on genital bulb (arrow in
Fig. 86
), and by internal female genitalia (
Figs 88
,
98
): anterior bilobed sclerite (arrows in
Fig. 88
); median arc between pore plates; pore plates far apart; note that female of
C. merida
Huber, 2000
is unknown.
Etymology
The species name refers to the
type
locality; noun in apposition.
Type
material
VENEZUELA
–
Yaracuy
•
♂
holotype
,
ZFMK
(Ar 21834),
Guaquira
, ‘site 1’ (
10.2951° N
,
68.6535° W
),
120 m
a.s.l.
,
forest along stream
,
16 Feb. 2020
(
B.A. Huber
,
O. Villarreal M.
,
Q. Arias C.
)
.
Other material examined
VENEZUELA
–
Yaracuy
•
3 ♂♂
,
6 ♀♀
,
ZFMK
(Ar 21835), and
8 ♀♀
in pure ethanol,
ZFMK
(Ven20- 158), same collection data as for holotype
•
1 ♂
,
6 ♀♀
,
ZFMK
(Ar 21836),
Guaquira
, ‘site 2’ (
10.2807° N
,
68.6530° W
),
150 m
a.s.l.
,
forest along stream
,
17 Feb. 2020
(
B.A. Huber
,
O. Villarreal M.
,
Q. Arias C.
)
.
Description
Male
(
holotype
)
MEASUREMENTS. Total body length 1.1, carapace width 0.55. Distance PME–PME
35 µm
; diameter PME
60 µm
; distance PME–ALE
30 µm
; AME absent. Leg 1: 8.45 (2.0 +0.2 +2.2+3.15 + 0.9), tibia 2: 1.1, tibia 3: 0.85, tibia 4: 1.1; tibia 1 L/d: 49.
COLOR (in ethanol). Prosoma and legs ochre-yellow, sternum whitish, legs without dark rings; abdomen pale greenish gray, dorsally with darker bluish marks.
Figs 85–88.
Canaima guaquira
Huber
sp. nov.
; from Yaracuy, Guaquira (type locality; ZFMK Ar 21835).
85–86
. Left male pedipalp, prolateral and retrolateral views (arrow: dorsal process on genital bulb).
87
. Male chelicerae, frontal view.
88
. Cleared female genitalia, dorsal view (arrows: anterior bilobed sclerite). Abbreviation: r =receptacle. Scale lines: 0.2 mm.
BODY. Habitus as in
Fig. 54
. Ocular area slightly raised. Carapace with distinct thoracic groove. Clypeus slightly more protruding than in female and with sclerotized rim. Sternum wider than long (0.40/0.30), with pair of small but distinct anterior processes. Abdomen oval.
CHELICERAE. As in
Fig. 87
, with very short entapophyses and three pairs of frontal processes; highly distinctive distal pair of slender curved processes.
PALPS. As in
Figs 85–86
; coxa with distinct retrolateral-ventral apophysis, trochanter with low ventral process, femur proximally with retrolateral process, distally with small ventral process (similar to
C. arima
; cf.
Huber 2000
: fig. 1322); procursus with distinctive dorso-distal sclerite and membranous elements; genital bulb with semitransparent dorsal process (arrow in
Fig. 86
), embolar process whitish, weakly sclerotized.
LEGS. Without spines and curved hairs; with higher than usual density of vertical hairs on tibiae; retrolateral trichobothrium of tibia 1 at 25%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~13 pseudosegments.
Male
(variation)
Tibia
1 in
five males (incl.
holotype
): 1.9–2.4 (mean 2.14).
Female
In general similar to male (
Fig. 55
) but without humps on sternum, clypeus less protruding and rim
not
sclerotized, few vertical hairs on leg tibiae. Tibia
1 in
13 females
: 1.50–1.75 (mean 1.65). Epigynum (
Fig. 96
) slightly protruding, anterior plate medially whitish, laterally brown, sclerotized; large light brown posterior plate. Internal genitalia (
Figs 88
,
97–98
) with anterior bilobed sclerite (arrows in
Fig. 88
), median arc between pore plates, pore plates far apart, and large anterior transparent receptacle.
Distribution
Known from
type
locality only, in
Venezuela
,
Yaracuy
(Fig. 1034).
Natural history
Most specimens were collected in dead bamboo sheaths on the ground but the species seemed to be common in the general forest leaf litter.