Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica Author Filippini, V. Author Galante, E. Author Micó, E. text The Coleopterists Bulletin 2015 2015-09-30 69 3 463 476 http://dx.doi.org/10.1649/0010-065x-69.3.463 journal article 10.1649/0010-065x-69.3.463 1938-4394 10106050 9A0330AD-EEF7-4A80-8D80-1B6116BD856F Anomala robiginosa Filippini, Galante, and Micó new species ( Figs. 6 , 14 , 22 , 30 , 34 ) Type material. Holotype : ♂ " Zarcero , Alfaro Ruiz , 1700m Alajuela Pr. COSTA RICA . Apr - May 1989 . A. Del Valle & A. Solis , L- N 240500_493500 / INBIOCRI002517719" deposited at MNCR . Paratypes (3): 1♀ " Santa Rosa National Park Guana. Prov. Costa Rica 18–20 May 1979 D. H. Janzen / INBIOCRI000111925" ; 1♂ " ALAJUELA , COSTA RICA Zarcero , 1700 m 23-V-87 COL: A. SOLIS B. / CEUA00106202 " ; 1♂ " Alajuela . Costa Rica . Zarcero , 1700 m 23 may 1987 Angel Solís / INBIOCRI002517717". Two paratypes are deposited at MNCR , and one paratype is deposited at Universidad de Alicante . Holotype . Male ( Fig. 6 ). Body oval. Length 11.23 mm , width 5.75 mm . Head, pronotum, and scutellum bronze-brown. Elytra ochre with suture and punctures reddish brown. Pygidium, legs, and venter orange-brown. Head: Clypeus trapezoidal, surface densely punctate-reticulate, convex. Anterior margin straight, thin in frontal view, width/ length ratio 2.09. Frons densely punctate, with a parabolic concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.48. Ratio of antennal funiculus/club 0.7. Pronotum: Trapezoidal. Ratio width/length 1.84. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, obliterated at middle. Surface with deep, dense punctures. Scutellum: Subtriangular, with straight lateral margins and acute apex. Ratio width/length 1.27. Surface densely punctate. Elytra: With costae defined by rows of sparse, coalescing punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Finely granulated, with long setae on posterior margin. Ratio width/length 1.65. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 3–4 rows of punctures per sternite. Last sternite punctate, with subapical bead thick and sinuate, apical membrane wide. Legs: Protibia ( Fig. 14 ) bidentate, apical tooth long and straight, basal tooth below internal apex of protibia triangular, sharp, obtuse. Metatibia slightly narrower sub- apically. Ratio length/width 2.98. First external carina with row of sparse setae. Surface punctate above second external carina and rugose below. Internal protarsal claw bifurcate, with upper branch slightly shorter and 1/3 width of lower one. Inferior margin with a sharp angle at half length. Male genitalia: Parameres slender ( Fig. 22 ), inferior margin sinuate. Apical portion of ventral plate membranous. Endophallus ( Fig. 30 ) developing dorsally with respect to median lobes; sclerotized, flat plate present posteriorly at base, above which a rounded, hairy inflation is present; ejaculatory duct opening located frontally. Female. Similar to male except protibia ( Fig. 14 ) with longer and wider first tooth; internal protarsal claw narrower with curve inferior margin; last sternite with curved apex. Variation. Three paratypes . Head, pronotum, and venter vary from orange to dark brown; presence of small dark maculae on basal third of elytra. Body length 10.68–11.49 mm , body width 5.75– 6.44 mm . Clypeal width/length ratio 2.01–2.33. Ratio of interocular width/width of eye 2.24–2.75. Ratio of antennal funiculus/club 0.67–0.74. Pronotal width/length ratio 1.72–1.84. Scutellar width/length ratio 1.24–1.37. Pygidial width/length ratio 1.65–1.82. Metatibial length/width ratio 2.77–3.03. Etymology. From the Latin adjective robiginosus, -a, -um , rusty, for the rust-colored elytral maculae. Distribution. Anomala robiginosa inhabits the dry forest in the northwestern Guanacaste lowlands and Tilarán and northwestern extreme of Central mountain ranges of Costa Rica , from 500 to 1,700 m elevation ( Fig. 34 ). Diagnosis. Pronotum bronze-brown, elytra with pigmented punctures, parameres slender, apex squared in lateral view, endophallus developing dorsally with respect to the median lobes. This species is very similar to A. sticticoptera , but it can be distinguished by the shape of the aedeagus. In A. sticticoptera , the parameres are shorter, with a narrow apex, and the ventral angle is acute and protruding posteriorly ( Fig. 33 ).