Systematics of the Protohermes costalis species-group (Megaloptera: Corydalidae) Author Liu, Xingyue Author Hayashi, Fumio Author Yang, Ding text Zootaxa 2007 1439 1 46 journal article 10.5281/zenodo.175948 02bb7dce-7590-4f45-8aa9-c6280f2e8b36 1175-5326 175948 Protohermes triangulatus sp. nov. ( Figs. 21 , 128–134 ) Diagnosis . Head with two small blackish markings on each side of vertex; pronotum with two large blackish markings on each side; male ninth sternum with deep V-shaped posterior incision; tenth tergum with an exceptionally well developed tufted tubercle at posteromedial corner. Body length 27–34 mm (male) and 35–41 mm (female); forewing length 37–41 mm (male) and 46–50 mm (female), hindwing length 32–36 mm (male) and 39–44 mm (female). Head ( Fig. 128 ) pale yellow without post-ocular spine; vertex with two small blackish markings on each side, lateral marking cuneiform, medial marking punctuate, and sometimes markings almost absent. Compound eyes brown; ocelli yellow, darkly margined medially. Antennae black, with scapes and pedicel yellow. Mouthparts yellow; mandible with distal 1/3 black. Occiput with one pair of blackish markings on lateral sides. Thorax yellow. Pronotum ( Fig. 128 ) with two small blackish markings on each side. Meso- and metanota each with one pair of brownish markings on lateral sides. Thoracic pilosity yellow, much longer on meso- and meta thorax. Legs yellow with dense yellowish short setae; tibiae and tarsi mostly dark, and sometimes only 2nd–5th mid and hind tarsomeres dark; tarsal claws reddish brown. Forewings ( Fig. 21 ) light smoky brown, with 1 whitish marking at base, 4–5 mostly fused whitish markings at middle and 1 round whitish marking on apical 1/3; costal cellules with distinct brownish stripes. Hindwings mostly hyaline, respectively with 1 indistinct round whitish marking on apical 1/3. Veins yellowish brown, with costal crossveins much paler. Rs 9- branched, last branch bifurcate or trifurcate; 9–10 crossveins between R1 and Rs; M1+2 4 to 6-branched, M3+4 2- branched; 1A 3-branched. FIGURES 128–134. Protohermes triangulatus , sp. nov. 128. Male head and pronotum, dorsal view. 129. Male genitalia, dorsal view. 130. Male genitalia, ventral view. 131. Male ninth gonostylus, caudal view. 132. Male tenth sternum, ventral view. 133. Female genitalia, lateral view. 134. Female eighth sternum, ventral view. Scale lines = 1.0 mm. Abdomen pale brown with yellowish short setae. Male ninth tergum ( Fig. 129 ) subquadrate, with arched anterior margin and shallowly incised posterior margin. Ninth sternum ( Fig. 130 ) broad with nearly straight lateral margin; posterior margin with deep V-shaped incision; central portion inflated. Ninth gonostylus ( Fig. 131) unguiform and slightly curved dorsomedially. Tenth tergum ( Figs. 129–130 ) short and subcylindrical, ventrally with one well developed tufted tubercle at posteromedial corner; posterolateral corner distinctly and acutely produced; tip slightly incised and densely setose. Tenth sternum ( Fig. 132 ) arched, anteriorly slightly elevated at middle; posterior incision V-shaped and moderately developed; lateral lobes digitiform and slightly incurved. Female eighth sternum ( Figs. 133–134 ) strongly sclerotized and subtrapezoidal in lateral view, with truncate posterior margin; ninth segment ( Fig. 133 ) laterally inflated in one pair of small suboval lobes; ninth gonocoxite valvate and membranous with one digitiform process at tip; tenth tergum short, with posterior margin medially incised, leaving one subtriangular upper and one round lower lobe. Material examined . Holotype ɗ, CHINA : Yunnan, Xinping, 19.VII.1980 ( CAU ). Paratypes Ψ, CHINA : Yunnan, Shuangjiang, 858 m , 5.VI.1980 , Wang Zhang ( CAU ); ɗ, CHINA : Yunnan, Yongping, 1700 m , 26.V.1980 ( CAU ); ɗ, CHINA : Yunnan, Anning, 2000 m , 17.VII.1980 ( CAU ); ɗ, CHINA : Yunnan, Fengqing, Songxiangchang, 1500 m , 3.VII.1980 , Fu Zhang ( CAU ); ɗ Ψ, CHINA : Yunnan, Gejiu, 11.VIII.1980 , Guoxiu Li ( CAU ); 1 without genitalia, CHINA : Yunnan, Yimen ( CAU ); ɗ, CHINA : Yunnan, Jingdong, 1170 m , 30.V.1956 ( IZCAS ); Ψ, CHINA : Yunnan, Jingdong, 1170–1300 m , 30.V.1956 ( IZCAS ); ɗ, CHINA : Yunnan, Jingdong, 1170 m , 2.VI.1956 ( IZCAS ); ɗ, CHINA : Yunnan, Lancang, 1000 m , 3.III.1957 , Shuyong Wang ( IZCAS ); Ψ, CHINA : Yunnan, Xishuangbanna, Yunjinghong, 2.VII.1958 , Chuanlong Li ( IZCAS ). Etymology . The specific epithet ‘ triangulatus ’ refers to the triangular or V-shaped posterior incision of the male ninth sternum in the new species. Distribution . China (Yunnan). Remarks . The new species appears to be closely related to P. arunachalensis , but can be easily separated by the V-shaped posterior incision of the male ninth sternum. In P. arunachalensis , the male ninth sternum has a U-shaped posterior incision.