The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications Author Sacchetti, Claudia 0000-0002-3225-3139 claudiasacc@icloud.com Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Ávila, Sérgio P. 0000-0002-3225-3139 claudiasacc@icloud.com text Zootaxa 2023 2023-05-24 5295 1 1 150 http://dx.doi.org/10.11646/zootaxa.5295.1.1 journal article 53396 10.11646/zootaxa.5295.1.1 82286fdc-a858-447c-9980-da2e8985d19c 1175-5326 7965273 F3A52660-70B8-439F-A7A0-F45ADC975EA5 Monoplex comptus (A. Adams, 1855 ) Plate 4 D 1 -D 2 * Triton comptus A. Adams 1855: 312 . Tritonium secans Bronn in Reiss 1862: 28 , pl, 1, fig.10. Tritonium secans Bronn — Mayer 1864: 72 , pl. 7, fig. 52. Triton ridleyi E. A. Smith 1890: 489 , pl. 30, fig. 1. Lampusia gracile (Reeve) — Dall 1889: 227 , pl. 29, fig. 2 ( non Triton gracilis Reeve, 1844 = Triton pfeifferianus Reeve, 1844). Cymatium comptum (A. Adams, 1855 ) — Yen 1942: 215 , pl. 18, fig. 112. Cymatium ( Septa ) gemmatum (Reeve, 1844) — Clench & Turner 1957: 222 , pl. 110, fig. 2, pl. 113, fig. 6, pl. 125, fig. 1-2 ( non Triton gemmatus Reeve, 1844 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Warmke & Abbott 1961: 101 , pl. 18b ( non Triton vespaceum Lamarck, 1822) . Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Kilias 1973: 193 , fig. 139 ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Abbott 1974: 163 , fig. 1754 ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Rios 1975: 79 , pl. 79, fig. 322 ( non Triton vespaceum Lamarck, 1822 ). Cymatium vespaceum ( Lamarck, 1822 ) — Hinton 1978: 30 , fig. 9 ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Kay 1979: 223 , fig. 79E (not 79F) ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Saunders 1980: 5 , upper fig. ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) vespaceum ( Lamarck, 1822 ) — Coelho et al. 1981: 124 , fig. 9 ( non Triton vespaceum Lamarck, 1822 ). Cymatium vespaceum ( Lamarck, 1822 ) — Garcia-Talavera 1983: 108 , pl. 5, fig. 2 ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Septa ) comptum . (A. Adams, 1855 )— Rios 1985: 75 , pl. 27, fig. 329. Cymatium ( Septa ) comptum (A. Adams, 1855 ) — Springsteen & Leobrera 1986: 113 , pl. 31, fig. 3a-b. Cymatium ( Turritriton ) comptum (A. Adams, 1855 ) — Garcia-Talavera 1987: 253 , pl. 2, fig. 5. Cymatium ( Turritriton ) comptum (A. Adams, 1855 ) — Lai 1989: 125 , fig. 44. Cymatium ( Turritriton ) comptum (A. Adams, 1855 ) — Henning & Hemmen 1993: 99 , pl. 21, fig. 5. Cymatium ( Septa ) comptum (A. Adams, 1855 ) — Rios 1994: 89 , pl. 29, fig. 345. Cymatium comptum (A. Adams, 1855 ) —Kubo, in Kubo & Kurozumi 1995: 75 , fig. 9. Cymatium vespaceum ( Lamarck, 1822 ) — Wellens & Wils 1997: 79 , fig. 14 ( non Triton vespaceum Lamarck, 1822 ). Cymatium ( Monoplex ) comptum (A. Adams, 1855 ) — Beu 1998b: 86 , fig. 23i, 26a-k. Cymatium ( Monoplex ) comptum (A. Adams, 1855 ) — Beu 1999: 21 , fig. 35. Cymatium ( Monoplex ) comptum (A. Adams, 1855 ) — Okutani 2000: 287 , pl. 142, fig. 11. Cymatium ( Turritriton ) comptum (A. Adams, 1855 ) — Redfern 2001: 62 , pl. 31, fig. 263A, B, pl. 109, fig. 263C. Cymatium comptum amphiatlanticum (A. Adams, 1855 ) — Garcia-Talavera & de Vera 2003: 202 , fig. 1-5. Cymatium ( Monoplex ) comptum (A. Adams, 1855 ) — Zhang & Ma 2004: 147 , text-fig. 91. Cymatium ( Monoplex ) comptum (A. Adams, 1855 ) —Beu & Segers in Poppe 2008: 644 , pl. 267, fig. 1, 3, 4, 7, 8. Monoplex comptus (A. Adams, 1855 ) — Beu 2010: 149 , pl. 36, figs. 10, 14. Cymatium comptum (A. Adams, 1855 ) —Hernández et al . 2011: 172, fig. 53L. Santa Maria material examined. Maximum height 25.0 mm, width 14 mm . Two fragments from Pinheiros, Santa Maria Island, Azores , Touril Complex, Lower Pliocene . Discussion. Tritonium secans Bronn, 1862 was based on four fragments, two from Pinheiros, one from Feteirinhas (= Ponta das Salinas ) and one from Cré. No further material is available, but the figure undoubtedly represents a Monoplex species , which in our opinion is M. comptum (A. Adams, 1855 ) . The specimen illustrated by Bronn in Reiss (1862 : pl, 1, fig.10; refigured pl. 4, fig. 4 D 1 ) and by Mayer (1864 : pl. 7, fig. 52; refigured pl. 4, fig. 4 D 2 ) is undoubtedly the same specimen, but the illustrator has highlighted the primary spiral cords much more strongly in Mayer’s illustration. For full discussion and comparison with related congeners, see Beu (2010: 149) . This species and Aspa marginata ( Gmelin, 1791 ) are the two amphiatlantic species present in the Azorean Pliocene assemblages. Distribution. Lower Pliocene: eastern Atlantic, Santa Maria Island, Azores (Bronn in Reiss 1862 ; Mayer 1864 ). Lower Pleistocene: eastern Atlantic, Moín formation, Costa Rica. Pleistocene or Holocene: Atlantic, Panama ; Red Sea , Egypt (NHMW coll.). Present-day: western Atlantic, Bermuda , Carolinian biogeographic province, Gulf of Mexico, Caribbean to Brasil ; eastern Atlantic, Canary Islands, Indian Ocean, and Red Sea (all references from Beu 2010 ).