Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous Author Szadziewski, Ryszard Author Dominiak, Patrycja Author Sontag, Elżbieta Author Krzemiński, Wiesław Author Wang, Bo Author Szwedo, Jacek text Zootaxa 2019 2019-10-24 4688 4 535 548 journal article 25149 10.11646/zootaxa.4688.4.5 c704c5d3-8ef6-48b6-a346-efe5b5e1c032 1175-5326 3517710 F412A472-E501-494B-9BF9-CAF253E9BD92 Culicoides myanmaricus Szadziewski & Dominiak, sp. nov. Figs 6 A–D urn:lsid:zoobank.org:act: 4671B5EB-F1B5-4F9C-830D-40E4965AA00A Diagnosis. This species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellomere 1 with sensilla coeloconica, flagellomeres 2–8 subcylindrical to cylindrical, gradually increasing in length distally; palpal segment 3 slender, greatly elongate, without sensory pit; proboscis moderately elongate; wing membrane with macrotrichia on distal 1/3. Male unknown. Description. Female ( Fig. 6A ). Head-body length minus antenna 1.15 mm . Proboscis ca. 0.25 mm long, 1.25 x longer than eye height. Eyes broadly separated ( Fig. 6B ). Antennal flagellum ( Fig. 6C ) ca. 0.39 mm long; first flagellomere with sensilla coeloconica; flagellomeres 2–13 gradually increasing in length; antennal ratio ca. 1.08. Palpus ( Fig. 6D ) 5-segmented, very slender, greatly elongate; total length 0.21 mm ; segment 3 greatly elongate, without sensory pit, length 0.10 mm . Wing ( Fig. 6A ) length 0.63 mm ; membrane with macrotrichia only on distal 1/3; basal radial cell without macrotrichia; costal ratio 0.81. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (3) 1.7. Abdominal sternite 8 with distinct caudomedian excavation. Male. Unknown. Material examined. Holotype female, NIGP 171165 (BA02 881). Burmese amber, Hukawng Valley , Myanmar . Syninclusion Hemiptera : Fulgoroidea of 3 rd or 4 th instar. Deposited in the collections of the Nanjing Institute of Ge- ology and Palaeontology , Chinese Academy of Sciences , Nanjing. Etymology. The species name refers to Myanmar ( Burma ) in Southeast Asia where Burmese amber is mined.