A range extension of a deep-sea barnacle of the genus Aurivillialepas (Cirripedia, Scalpellomorpha), a Macaronesian and amphitropical refugial genus having Mesozoic affinities
Author
Innocenti, Gianna
Author
Geronimo, Raffaella Di
Author
Newman, William A.
text
Zootaxa
2015
3974
2
257
266
journal article
10.11646/zootaxa.3974.2.9
7bd4550d-20b4-4ea7-9521-af7168ffd0f6
1175-5326
239470
4BE3EA89-1864-43E9-B813-320696C62958
Family
Calanticidae
Zevina, 1978
Subfamily
Calanticinae
Zevina, 1978
(
nom. trans.
Newman 1996
)
Calantica
Gray, 1825
.
Type
Scalpellum villosum
Leach, 1824
(cf.
Newman & Jones 2011
) Subfamily Scillaelepadinae Innocenti,
Di
Geronimo and Newman, nov.
Scillaelepas
Seguenza, 1872
.
Type
†
Pollicipes carinata
Philippi, 1835
Gruvelialepas
Newman, 1980
.
Type
Scalpellum pilsbryi
Gruvel, 1911
Aurivillialepas
Newman, 1980
.
Type
Scalpellum calyculus
Aurivillius, 1898
Newmanilepas
Zevina & Yakhontova, 1987
(
incertae sedis
).
Type
Scillaelepas mirafica
Zevina, 1976
Definition.
Calanticidae
with peduncular plates relatively large, covered by an inconspicuous cuticle, capitular plates basically thirteen in number in
Scillaelepas
(
Newman & Jones, 2011
)
. The plates consist of the unpaired rostrum (R), carina (C) and subcarina (SC), and paired rostrolatus (RL), scutum (S), lateral (L), tergum (T) and carinolatus (CL), supplemented with one subrostrum (SR) in
Aurivillialepas
Newman, 1980
, or two subrostrolatera (SR1 and SR2) in
Gruvelialepas
Newman, 1980
. The genus
Newmanilepas
Zevina & Yakhontova, 1987
, represented by
Scillaelepas mirafica
Zevina, 1976
, noted as atypical when first described (
Newman 1980
), subsequently proved to be an early ontogenetic stage to which a second pair of CLs and a SC are added (
Zevina & Yakhontova 1987
). Therefore it has been designated an
incertae sedis
member of the Scillaelepadinae.
Remarks.
Originally, most of the species comprising these four genera were assigned to
Scillaelepas
s.l.
and while new species have since been attributed to
Scillaelepas
s.s.
and
Aurivillialepas
, there has been no indication they do not form a natural group. Therefore, we recognized them as such. Nonetheless, while the calanticids already consists of two very large, one moderate and three small subfamilies, all but the Pollicipedinae and the new subfamily are in need of thorough revisions. This is especially true of the
Calanticinae
since what is now known of thoracican genetics (
Herrera
et al
. 2015
), the
type
species,
Calantica villosa
(
Leach, 1824
)
is among the few species rooting the only wholly scalpellomorph clade. This can be best appreciated when the unsupported nodes of their “maximum clade credibility ultrametric timescaled tree” are collapsed. Unfortunately, to date but a few calanticids and no scillaelepadines have been sequenced.