Prodromus of a fern flora for Bolivia. XXXV. Dryopteridaceae Author Kessler, Michael Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland Author Moran, Robbin C. New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA Author Mickel, John T. New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA Author Matos, Fernando B. Universidade Federal do Paraná, Departamento de Botânica, Caixa Postal 19031, 81531 - 980, Curitiba, PR, Brasil Author Smith, Alan R. Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland & University Herbarium, 1001 Valley Life Sciences Bldg. # 2465, University of California, Berkeley, CA 94720 - 2465, USA & Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland text Phytotaxa 2018 2018-06-11 353 1 448 450 http://dx.doi.org/10.11646/phytotaxa.353.1.1 journal article 10.11646/phytotaxa.353.1.1 1179-3163 Polystichum rufum M.Kessler & A.R.Sm., Brittonia 57(3): 217, f. 2d, 4. 2005. Range: — Peru and Bolivia (CO, LP, SC); likely to occur further north in the Andes. Ecology: —Common; terrestrial in humid forests, along forest margins, and in plantations; 700–1800 m . Notes: —Differs from P. montevidense in having uniformly orange large petiolar scales, and in growing at lower elevations ( 700–1800 m vs . 2100–4050 m ). Overall, a more delicate species with thinner-textured pinnules. Some specimens of P. rufum growing at especially low elevations ( 700 m ) approach P . albomarginatum in having dark-centered large petiolar scales, even though these are not sharply bicolorous. McHenry & Barrington (2014) included a single specimen, from Bolivia , in their analysis.