A revision of Phlegoninae (Coleoptera: Eucnemidae), with descriptions of a new genus and four new species Author Otto, Robert L. text Insecta Mundi 2017 2017-08-25 2017 569 1 27 journal article 10.5281/zenodo.5169054 1942-1354 5169054 5265BC7E-7076-4225-B659-678E3990B42C Phlegon giganteus Fleutiaux, 1929: 36–37 Fig. 9–14 Diagnosis. Oblong punctures along elytral sutures distinguish this species from P. buqueti , P. herculeanus and P. panamensis . Shape of pronotum further distinguishes female P. giganteus from P. chiriquiensis , as it gradually narrows anteriorly in P. giganteus and is parallel-sided in P. chiriquiensis . Specimens Examined. Six specimens were available for study: COSTA RICA : 1, “C. RICA: Gte prov., 1 mi. E. Los Angeles, 1-25-74, W. Vandevender.” / “J. Muona det., Phlegon , giganteus , Fleut. ” (genus, species and author handwritten) / “ Phlegon giganteus , Fleutiaux 1929 , J. Muona det. 2014” (JMC); 1, “CR. Punt. Penin de OSA, Los Banzos de Rincon, 60m , 30-12-95, L. Aryubo, LS985200N/515600E” (illegible handwritten label) / “ Phlegon giganteus , Fleutiaux 1929 , D. Alaruikka det. 2007, J. Muona conf. 2015” (INBC); 1, “Casa Oeste, Cerro El, Hacha, 12km SE La Cruz, 300m , Guanacaste Prov. , COSTA RICA, Jan 1988 , A. Solis” / “ Phlegon giganteus , Fleutiaux 1929 , D. Alaruikka det. 2007, J. Muona conf. 2015” (INBC); 1, “Casa Oeste, Cerro El, Hacha, 12km SE La Cruz, 300m , Guanacaste Prov. , COSTA RICA, Jan 1988 , M. Espinosa” / “ Phlegon giganteus , Fleutiaux 1929 , D. Alaruikka det. 2007, J. Muona conf. 2015” (INBC); 1, “Los Almendros, P.N., Guanacaste , Prov. Guan., acaste, Costa Rica, 28, mar a 24 abr 1992 , M. Reyes, L-N 334800, 369800” / “ Phlegon giganteus , Fleutiaux 1929 , D. Alaruikka det. 2007, J. Muona conf. 2015” (INBC); 1, “COSTA RICA Guan., 3km SE R. Naranjo, 1–10 Apr. 1992 , F.D. Parker” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green framed, white label) / “ Phlegon , giganteus , Fleutiaux, Det. R.L. Otto, 2005 ” (GERP). Redescription. ♂ : length 18.0–20.0 mm, width, 6.5–7.5 mm . ♀ : length 25.0–31.0 mm, width, 8.0–9.0 mm. Body stout, oblong, elongate; uniformly dark brown; antennae dark brown; femora, tibiae and tarsi dark brown; head, pronotum and elytra clothed with short, semierect copper-colored setae ( Fig. 9–10 ). Head: Subspherical; frons depressed above frontoclypeal region; surfaces shiny, densely punctate to almost confluently rugose; apical margin of frontoclypeal region rounded, about 2 times wider than base; mandibles slender, without teeth. Antenna: ♂ ( Fig. 11 ): About 3/4 of body length; antennomere III slightly longer than IV; antennomeres IV–VIII quadrate, subequal; antennomeres IX–XI enlarged, tubular, dentate; antennomere X slightly shorter than either IX or XI; antennomeres IX and XI each as long as III–VIII combined; antennomere X as long as VI–VIII combined. ♀ ( Fig. 12 ): About 1/2 of body length; antennomere III slightly longer than IV; antennomere IV slightly longer than wide; antennomeres V–VIII quadrate, subequal; antennomeres IX–XI enlarged, tubular, very weakly dentate; antennomere IX slightly longer than either X or XI; antennomere IX as long as VI–VIII combined; antennomeres X and XI each slightly longer than VI–VIII combined. Pronotum: Surfaces shiny, punctures closely spaced; slightly wider than long, with large, sharp hind angles; sides anteriorly narrower, slightly constricted basally above hind angles; disc convex, variably with four circular foveae, anterior pair much more impressed; base sinuous. Scutellum: Setose, quadrate, elongate and distally rounded. Elytra: Striae indicated, formed from elongate punctures; interstices slightly elevated; surfaces shiny, shallowly punctate to slightly rugose; apices ( Fig. 13 ) with shallow, V-shaped cavities near elytral suture. Legs: First tarsomere shorter than the combined length of remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV excavated, bilobed; metatarsomere V elongate; pretarsal claws simple. Venter: Closely punctate, with short, semierect to recumbent copper-colored setae; hypomeron simple, without lateral antennal grooves; metepisterna parallel-sided; metacoxal plates medially 3.0–6.0 times wider than laterally. Aedeagus ( Fig. 14 ): Basal piece short, rounded, dorsally open; remaining parts bulbous, basally narrowed, widest near base of lateral lobes; median lobe short, ventrally curved, dorsally hooked at tip; lateral lobes ventrally curved, elongate, apically rounded and divergent. Distribution. Taken from five localities throughout much of Costa Rica . Biology. Some adults have been taken from montane cloud forests. Larvae and pupae are unknown. Note. Muona (1985) reported finding the species at Izabal , Rio Dulce in Guatemala , but I was unable to examine and verify the identity of the eucnemid collected in Guatemala during the course of this study. The types of this species collected in Turrialba, Costa Rica , were not available for study.