A Rosetta Stone for eastern Pacific Caecidae (Gastropoda: Caenogastropoda) Author Raines, Bret K. text Zootaxa 2020 2020-08-11 4827 1 1 146 journal article 8690 10.11646/zootaxa.4827.1.1 c93b6c76-aaac-4fcd-acca-3dc3e0fd62d4 1175-5326 4402308 2CE528C2-DE07-4155-B37D-BE2C342A3901 Caecum magellanicum (di Geronimo, Privitera & Valdovinos, 1995 ) Figures 96 A–E, 97A–F, 98A–B, 126F Fartulum magellanicum di Geronimo et al . 1995: 114 , figs. 1–5, holotype IPOP M1.8.8.95, Cabo Deseado , Strait of Magellan , Chile . Fartulum magellanicum — Linse 2002: 85 , pl. VII, figs. 54–56; Mattano & Zelaya 2013: 170 . Material examined. In addition to the type material the following lots were examined. NHMLAC: LACM 1973 - 75.4 , Westhoff Island , Los Lagos Prov. , Chile , 23 m , 8 sh; LACM 1973 - 74.5 , Darwin Channel , Aysen , Prov., Chile , 8 m , 5 sh. FS: Stanley , near Yorke Point , East Falkland , Falkland Islands , 9 m . Original diagnosis. “Teleoconch cylindrical, moderately curved, vitreous or lightly opaque in less fresh shells. Under the optical microscope, surface seems entirely smooth, with less distinct annular growth striae. Using the SEM these striae are rather thick, irregularly distanced and little engraved. Apertural end circular with a thin lip folded outwards. Septum well developed, oblique angled to the back of the shell. Maximum diameter, at aperture, is constant to middle of the teleoconch while the posterior diameter is always smaller. Operculum and soft parts not known.” di Geronimo et al. (1995: 114) . FIGURE 96. Caecum magellanicum . A . Profile view (LACM 1973-75.4). B . Profile view (LACM 1973-75.4). C . Ventral view (LACM 1973-75.4). D . Operculum, exterior surface (LACM 1973-75.4). E . Holotype, IPOP M1.8.8.95. Description. Protoconch paucispiral (about 1.4 whorls), planispiral, smooth ( Fig. 97A ), transition to teleoconch I marked by very slight axial line ( Fig. 97A ). Teleoconchs I and II short, no sculpture observed, transition to teleoconch III marked by very slight axial line ( Fig. 97A ). Teleoconch III sculpture smooth with occasional axial growth striae ( Fig. 97C ), transition to teleoconch IV marked by incised axial line ( Fig. 97B ). Teleoconch IV with identical characters as previous stage, transition to teleoconch V marked by incised axial line ( Fig. 97B ). Teleoconch V (last growth stage) small for genus [Tol 1.26–1.45 mm ], tubular, thin, very narrow, mildly to moderately and regularly arched [Larc: 0.45–0.55 mm ; Arc: 0.13–0.14 mm ], subcylindrical, semi-translucent to opaque white ( Figs. 96 A–B). Surface smooth with numerous weak annuli ( Fig. 97E ). Posterior [Dpe: 0.20–0.22 mm ] with squared shoulder ( Fig. 97D ). Septum mucronate ( Fig. 97D ). Mucro heel-like, positioned along dorsal margin, usually covered by septum in adult stage ( Fig. 97D ). Aperture [Da: 0.24–0.25 mm ] thin ( Fig. 97F ). No varix. Lip smooth, with deflected peristome ( Fig. 97F ). Periostracum not observed. Operculum thin, light brown; exterior surface with 5–6 concentric rings ( Fig. 97D ). Conceptual reconstruction of growth stages shown in Fig. 98A . Distribution and Habitat. From Westhoff Island, Chile to Stanley, East Falkland, Falkland Islands . Subtidal, 8– 245 m . Uncommon on organogenic sand substrates. Remarks. The type material of Caecum magellanicum is comprised of three late subadult stages and nine adult specimens ( holotype IPOP M1.8.8.95, Fig. 96E ). Caecum magellanicum is the southern-most caecid known in the eastern Pacific and has also been found in the southern Atlantic. FIGURE 97. Caecum magellanicum shells at different growth stages. A . Protoconch and teleoconchs I to III (paratype, IPOP unnumbered). B . Teleoconchs III to V (LACM 1973-75.4). C . Surface of teleoconch IV (LACM 1973-75.4). D . Teleoconch V (LACM 1973-75.4). E . Surface of teleoconch V (LACM 1973-75.4). F . Close-up of aperture (LACM 1973-75.4). FIGURE 98. Caecum magellanicum . A . Conceptual reconstruction of growth stages. B . Species distribution map. Despite repeated attempts to obtain better images, the figures shown of the type material ( Figs. 96E ; 97A ) are from di Geronimo et al. (1995) . Hence it was not possible to observe the sculpture of teleoconchs I and II.