Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae)
Author
Luo, Jiu-Yang
C9ACC9F1-9C82-43C1-A463-5FCAA68557E9
State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China.
luojy93@mail.sysu.edu.cn
Author
Xie, Qiang
ABD76A07-9958-497F-B84E-03E86DBE8CCE
State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China.
xieq8@mail.sysu.edu.cn
text
European Journal of Taxonomy
2022
2022-03-11
802
1
57
http://dx.doi.org/10.5852/ejt.2022.802.1687
journal article
20211
10.5852/ejt.2022.802.1687
f446314a-e9b6-4217-a8fd-bb24423e54ca
2118-9773
6358878
C3F3CDB9-0703-458A-BCE4-DE81614F075B
Key to the species of
Kokeshia
Miyamoto, 1960
1. Tergite VIII not subdivided into two hemitergites, only a long,flattened, acuminate process on the right portion of tergite VIII, bent at about ⅓ from its apex (
Fig. 28A
) (
China
) .....
K. acutiformis
sp. nov.
– Tergite VIII subdivided into left and right hemitergites ................................................................... 2
Fig. 25.
Kokeshia zhengi
Rédei, Ren & Bu, 2012
.
A‒D
. Holotype, ♂ (NKUM).
A
. Dorsal view.
B
. Lateral view.
C
. Pregenital abdomen and genitalia, dorsal view.
D
. Labels.
2. Process of left hemitergite VIII elongated, needle-shaped, flagelliform or filiform ......................... 3
– Process of left hemitergite VIII not needle-shaped, flagelliform or filiform .................................... 6
3. Process of left hemitergite VIII significantly long, almost reaching the width of tergite VII, the process with an appendage split into three to five branches (
Fig. 17D
;
Ren & Zheng 1992: 191
, figs 21‒23) (
China
) ....................................................................................
K. hsiaoi
Ren & Zheng, 1992
– Process of left hemitergite VIII relatively long, more than half the width of tergite VII, the process without any appendage ..................................................................................................................... 4
Fig. 26.
Forewings of
Kokeshia
spp.
, dorsal views.
A
.
K. acutiformis
sp. nov.
, paratype, ♂ (SYSBM).
B
.
K. baii
sp. nov.
, paratype, ♂ (SYSBM).
C
.
K. bui
sp. nov.
, paratype, ♂ (SYSBM).
D
.
K. caii
sp. nov.
, paratype, ♂ (SYSBM).
E
.
K. drepanoides
sp. nov.
, paratype, ♂ (SYSBM).
F
.
K. hilli
sp. nov.
, paratype, ♂ (SYSBM).
G
.
K. hsiaoi
Ren & Zheng, 1992
, ♂ (SYSBM).
H
.
K. pengae
sp. nov.
, paratype, ♂ (SYSBM).
4. Process of left hemitergite VIII filiform, rectangularly bent, with spiniform microtrichia on its apex (
Štys 1985: 200
, figs 37–38, 40) (
Nepal
) ...........................................................
K. similis
Štys, 1985
– Process of left hemitergite VIII needle-shaped or flagelliform, not filiform, without spiniform microtrichia on its apex .................................................................................................................... 5
5 Process of left hemitergite VIII slender, flagelliform, curved from approximal middle portion (
Rédei 2008: 244
, fig. 6; 248, fig. 14) (
Thailand
) .......................................................
K. oroszi
Rédei, 2008
_ Process of left hemitergite VIII slender, needle-shaped, curved from base (
Fig. 29D
) (
China
) ...................................................................................................................
K. pengae
sp. nov.
6. Process of left hemitergite VIII with a broad brush-like structure formed by hair-like setae .......... 7
– Process of left hemitergite VIII without broad brush-like structures formed by hair-like setae ...... 8
7. Brush-like structure on left hemitergite VIII formed by several narrow, hair-like setae; apical portion of phallus forming about one incomplete coil outside of pygophore (
Štys 1985: 192
, figs 15, 17; 194, figs 19–20; 196, fig. 26) (
Nepal
) ....................................................................
K. martensi
Štys, 1985
– Brush-like structure on left hemitergite VIII formed by less numerous and distinctly thicker setae; apical portion of phallus forming more than a half coil outside of pygophore (
Rédei 2008: 244
, fig. 5; 246, fig. 7) (
India
......................................................................................
K. stysi
Rédei, 2008
8. Process of left hemitergite VIII triangular or finger-shaped ............................................................. 9
– Process of left hemitergite VIII not triangular or finger-shaped ..................................................... 14
9. Process of left hemitergite VIII triangular or subtriangular ............................................................ 10
– Process of left hemitergite VIII finger-shaped ................................................................................ 12
Fig. 27.
Forewings of
Kokeshia
spp.
, dorsal view.
A
.
K. redeii
sp. nov.
, paratype, ♂ (SYSBM).
B
.
K. renae
sp. nov.
, paratype, ♂ (SYSBM).
C
.
K. weirauchae
sp. nov.
, paratype, ♂ (SYSBM).
D
.
K. xiei
Rédei, Ren & Bu, 2012
. ♂ (SYSBM).
10. Process of left hemitergite VIII with a broad, thick near triangular process, with about 8 thick spiniform microtrichia at its right posterolateral margin (
Rédei et al. 2012: 30
, fig. 2; 31, fig. 4; 33, figs 8‒10) (
China
) .........................................................................
K. zhengi
Rédei, Ren & Bu, 2012
– Process of left hemitergite VIII near triangular, not broad ..............................................................11
11. Process of left hemitergite VIII elongated triangular, right margin with about nine, thick, nearly parallel spiniform microtrichia (
Fig. 28C
) (
China
) .....................................................
K. bui
sp. nov.
– Process of left hemitergite VIII subtriangular, without thick spiniform microtrichia (
Fig. 30B
) (
China
) .....................................................................................................................
K. renae
sp. nov.
12. Process of left hemitergite VIII elongate, curved, finger-shaped, with one in apex, and two more thick spiniform microtrichia in ventral side of distal portion, right hemitergite VIII with lamellate process above tergite VII (
Fig. 28B
) (
China
) .............................................................
K. baii
sp. nov.
– Process of left hemitergite VIII moderate long, finger-shaped, right hemitergite VIII without lamellate process above tergite VII ................................................................................................. 13
13. Process of left hemitergite VIII curved, finger-shaped, with about six thick spiniform microtrichia at ventral side of apex; apical portion of phallus forming about two and a half coils outside of pygophore (
Fig. 30C
) (
China
) .......................................................................
K. weirauchae
sp. nov.
– Process of left hemitergite VIII not curved, finger-shaped, with out thick spiniform setae;apical portion of phallus forming incomplete one coil outside of pygophore (
Fig. 28D
) (
China
) ....
K. caii
sp. nov.
14. Process of left hemitergite VIII long, thick and horizontally directed, with numerous thick spiniform microtrichia at its ventral surface; apical portion of phallus forming about one coil outside of pygophore (
Fig. 30D
;
Rédei et al. 2012: 30
, fig. 1; 31, fig. 3; 33, figs 5‒7) (
China
) ...............................................................................................
K. xiei
Rédei, Ren & Bu, 2012
– Process of left hemitergite VIII not as above .................................................................................. 15
15. Process of left hemitergite VIII short and thick, hook-shaped, rectangularly bent; apical portion of phallus forming about one and a half coils outside of pygophore (
Miyamoto 1960: 171
, pl. 19h–i) (
Japan
) .......................................................................................................
K. esakii
Miyamoto, 1960
– Process of left hemitergite VIII not as above .................................................................................. 16
16. Process of left hemitergite VIII flattened, broad and curved, sickle-shaped; apical portion of phallus forming about a half coil outside of pygophore (
Fig. 29A
) (
China
) ............
K. drepanoides
sp. nov.
– Process of left hemitergite VIII not as above .................................................................................. 17
17. Process of left hemitergite VIII flattened, slender and curved, sword-shaped; apical portion of phallus forming about one coil outside of pygophore (
Fig. 30A
) (
China
) ...........................
K. redeii
sp. nov.
– Process of left hemitergite VIII not as above .................................................................................. 18
18. Process of left hemitergite VIII nearly quadrangular, left portion with a raised projection, which bearing about seven thick basal-curved spiniform microtrichia, posterior margin of the process with about seven thick basal-curved spiniform microtrichia; apical portion of phallus forming more than one coil outside of pygophore (
Fig. 29B
) (
China
) ......................................................
K. hilli
sp. nov.