Three new synonyms within the flower chafer genus Goliathopsis Janson, 1881 (Coleoptera: Scarabaeidae: Cetoniinae) from China
Author
Xu, Hao
Author
Qiu, Jian-Yue
text
Zootaxa
2020
2020-06-08
4789
1
91
131
journal article
10.11646/zootaxa.4789.1.3
1175-5326
3884725
CA5C0D09-B499-4077-873F-BF5B55C504B2
Goliathopsis cervus
Janson, 1881
(
Figs. 5
,
13
,
22–23
,
52–57
,
89–90
,
103–108
,
164
)
Goliathopsis cervus
Janson, 1881: 610
(type locality:
Siam
[=
Thailand
]), plate XI, fig.
4 ♂
,
5 ♀
;
Bergé 1884: 158
;
Gestro 1888: 119
, fig.
♂
(horn);
Fairmaire 1893b: 316
;
Arrow 1910: 206
(synonymized with
G. despectus
);
Janson 1917: 110
(status revalidated);
Schenkling 1921: 364
(in synonymy
G. despectus
);
Krikken 1977: 314
(
lectotype
designated);
Antoine 1991: 9
(status resurrected);
Krajčík 1999: 39
;
Krajčík 2012: 116
.
Type material.
As
Janson (1881)
stated in the original publication, this species was originally described from
Thailand
based on a pair of specimens. The type specimens now are preserved in
RMNH
, and the male was designated as
lectotype
by
Krikken (1977)
. Due to
RMNH
is still closed for collection integration (Jan Krikken, personal com-
munication, 2019), the female
paralectotype
is unavailable for the present work, and the male
lectotype
was simply examined by photographs (
Figs. 103–108
):
Goliathopsis cervus, Jans.
,
Type
,
Siam
//
♂
TYPE
.
FIGURES 13–21.
Cephalic horns of
Goliathopsis
Janson, 1881
species (dorsal view). 13,
G. cervus
Janson, 1881
; 14,
G. despectus
(
Westwood, 1873
)
, paralectotype of
G. capreolus
Gestro, 1888
in MNHN; 15,
G. duponti
Antoine, 1991
; 16,
G. es- quiroli
Pouillaude, 1913
; 17–18,
G. ferreroi
Antoine, 1991
; 19,
G. gressitti
Ruter, 1978
; 20,
G. lameyi
Fairmaire, 1893
; 21,
G. velutinus
Pouillaude, 1913
.
Other material examined
(
2♂♂
,
1♀♀
).
THAILAND
:
2♂♂
(
KSCJ
),
Khao Bandai It
,
Petchaburi Prov.
C. Thailand
,
2012.V.22
,
S.Tsuyuki
leg. //
Goliathopsis velutinus
Pouillaude, 1913
, det.
K. Sakai
, 2015 [misidentification]
;
1♀
(
KSCJ
),
Khao Soi Dao
,
Chanthaburi
,
Thailand
,
2012.V.17–21
,
K.Takahashi
leg. //
Goliathopsis velutinus
Pouillaude, 1913
, det.
K. Sakai
, 2015 [misidentification]
.
Diagnosis.
Smallest species of the genus, body length 9.5–10.5 mm; width 5.3–5.5 mm. Dorsal surface and pygidium clad with brown tomentum (
Figs. 54, 56
,
103, 106
). Midline on pronotum and scutellum yellow, distinct. Elytron usually with two light brown maculae (not distinct): a small one on posthumeral area, sometimes absent; the other small one on the mediodiscal area near sutural costa. Disc of elytron with U-shaped (few drop-shaped), setiferous punctures (
Fig. 5
); setae short. Pygidium with small, rounded, setiferous punctures (
Figs. 22–23
). Cephalic horns of male small and slender, inner side with tomentum, not constricted at base and not distinctly expanded at apex (
Figs. 13
,
103
). Male genitalia small (
Figs. 52–53
), proximal and distal parts of parameres gradually expanded in apical view, medially constricted, apex rounded and inner side slightly curved; parameres slightly curved in lateral view, basal piece broad.
FIGURES 22–37.
Pygidium of
Goliathopsis
Janson, 1881
species (male in left, female in right; posterior view). 22–23,
G. cer- vus
Janson, 1881
; 24–25,
G. despectus
(
Westwood, 1873
)
, lectotype and paralectotype of
G. capreolus
Gestro, 1888
in MSNG; 26–27,
G. duponti
Antoine, 1991
; 28–29,
G. esquiroli
Pouillaude, 1913
; 30–31,
G. ferreroi
Antoine, 1991
; 32–33,
G. gressitti
Ruter, 1978
; 34–35,
G. lameyi
Fairmaire, 1893
; 36–37,
G. velutinus
Pouillaude, 1913
.
This species is closed to
G. velutinus
, but body size distinctly smaller (
Figs. 89–90
); cephalic horns smaller and not constricted at base (
Fig. 13
; larger and constricted at base in
G. velutinus
,
Fig. 21
); punctures on elytral disc usually U-shaped (
Fig. 5
; usually drop-shaped in
G. velutinus
,
Fig. 12
); parameres smaller and distal parts gradually expanded (
Fig. 52
; larger and distinctly expanded in
G. velutinus
,
Fig. 42
).
Distribution.
Thailand
.
Remarks.
Based on the good illustrations of
Goliathopsis cervus
in
Janson (1881)
,
Gestro (1888)
pointed out several differences between
G. cervus
and his new species
G. capreolus
, especially in the shape of the cephalic horns. Nevertheless,
Arrow (1910)
regarded them as the same species and listed both as synonyms of
G. despectus
. In response to Arrow’s treatment,
Janson (1917)
insisted on the validity of
G. cervus
and provided more differences between females based on a re-examination of the female type of
G. despectus
in OXUM. Unfortunately, this disputation was not mentioned by subsequent authors. In the most recent work,
Antoine (1991)
also treated
G. cervus
as valid species mainly by the difference of the tomentose area on cephalic horns and elytra according to the figures in
Janson (1881)
. By examining the fresh specimens collected from southern
Thailand
and the male
lectotype
, several distinct differences from
G. despectus
are observed: cephalic horns of male smaller and inner side with tomentum in
G. cervus
(
Fig. 13
; larger and glabrous in
G. despectus
,
Fig. 14
); pygidium subtriangular in posterior view in
G. cervus
(
Figs. 22–23
; oval in
G. despectus
,
Figs. 24–25
); parameres smaller and apex rounded in
G. cervus
(
Fig. 52
; larger and apex sharp in
G. despectus
,
Fig. 48
). Consequently,
Goliathopsis cervus
is an independent species.