Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records Author Zorn, Carsten Author Kobayashi, Hirokazu Author Wada, Kaoru text Beiträge Zur Entomologie = Contributions to Entomology 2017 67 2 325 352 journal article 10.21248/contrib.entomol.67.2.325-352 0005-805X 5742448 F2CE9642-9537-46FF-BA3F-5D4148AC1A0B Anomala imperspicabilis spec. nov. urn:lsid:zoobank. org:act: 8B865ECA-A674-4C60-A04A-10B0B4196C34 ( Figs 4 , 13 ) Type series : Holotype : , VIETNAM Nghe An prov. Con Cuong JUN. 2008 . ( NSMT ) . Paratypes : 1 , the same data as for the holotype ( MFPC ) ; 1 , 1 , LAOS-NE; HUA PHAN prov. BAN SALUEI; Phu Phan Mt. ; 20°15'N 104°02'E ; 1500-2000 m ; 26.iv.-11.v.2001 J. Bezděk & D. Hauck leg. ( CZPC ) ; 9 , 24 , NE-LAOS: Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) ~ 20°12'N , 104°01'E ; 1300-1500 m ; 01.-31.V.2011 ; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ( CZPC , PLPC , ZFMK ) ; 28 , 25 , NE-LAOS: Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. )- 20°12'N , 104°01'E 11.iv.-15.v.2012 , 1300-1900 m leg. C. Holzschuh – ZFMK Ankauf 2012 ( CZPC , ZFMK ) ; 3 , 5 , NE-Laos: Hua Phan Prov. , Ban Saleui , Phou Pan ( Mt. ) - 20°12'N , 104°01'E , 11.iv.-15.v.2012 , 1300-1900 m , leg. C. Holzschuh ZFMK Ankauf 2012/13 ( CZPC , ZFMK ) ; 5 , 1 , NE-LAOS Hua Phan prov. Ban Saleui Phou Pan ( Mt. ) 20°12'N , 104°01'E 11.iv.-15.v.2012 , 1300-1900 m leg. C. Holzschuh ZFMK Ankauf 2012/13 ( CZPC , ZFMK ) ; 3 , 3 , NE-LAOS: Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) ~ 20°12'N , 104°01'E ; 1300-1900 m leg. C. Holzschuh 11.iv.-15.v.2012 – ZFMK Ankauf 2012/13 ( CZPC , ZFMK ) ; 7 , 5 , LAO-NE Hua Phan Prov. , ~ 20°12'N 104°01'E , PHU PHAN Mt. , 1500-1900 m , 17.v.-3.vi.2007 , Vit Kubáň leg. ( NMPC ) ; 28 , 17 , LAOS-NE, Houa Phan prov. , 20°13'09-19"N 103°59'54"- 104°00'03"E , 1480- 1510 m , PHOU PANE Mt. , 22.iv.-14.v.2008 Vit Kubáň leg. ( CZPC , NMPC ) ; 1 , 1 , LAOS-NE, Houa Phan prov. , 20°12-13.5'N 103°59.5- 104°01'E , Ban Saleui-> Phou Pane Mt. , 1340-1870 m , 15.IV.-15.V.2008 , lao collectors leg. ( NMPC ) . Description : Body shape elongate ovoid, strongly convex; body length: 15.3-18.6 mm , width: 8.5-9.4 mm . Color . Dorsal face metallic green, pronotum with vague narrow yellow border (lateral margin of pronotum and margins of elytra broadly suffused with yellow color in specimens from northern Laos ); footstalk of antenna yellowish brown, club black; ventral face predominantly metallic green; mouthparts, sternal segments, femora and tibiae yellowish brown, meso- and metatibia with dark metallic green distal ends; tarsi dark metallic green. Head with erect yellow setae ( 0.13-0.25 mm in length) along eyes; interocular distance equals 0.74 times the maximum transverse head width. Antennal club 0.8 times as long as footstalk. Clypeus broadly trapezoidal with broadly rounded front angles; densely punctate, the punctures round and partly coalescent; anterior margin distinctly reflexed; fronto-clypeal suture almost straight. Frons densely punctate, partly coalescent and rugulose in lateral portions, the punctures becoming larger and sparser postero-laterad; vertex irregularly scattered with smaller round punctures. Pronotum 1.9 times as wide as long; sides broadly rounded, widest shortly before the base; basal marginal line almost complete, interrupted before the scutellum; front angles produced and acute, hind angles obtuse; disc with dense transverse punctures (ca. 0.05-0.1 mm ); punctures partly transversely confluent towards front- and hind angles; with a distinct longitudinal medial furrow and vague depression in the middle of the lateral area; with erect yellow setae ( 0.25-0.38 mm in length) along lateral margins. Scutellum 1.3 times as wide as long, apex acute; irregularly scattered with somewhat transverse punctures. Elytra . Regularly striate-sulcate; intervals moderately convex; striae not marked by regular rows of punctures; entire surface covered with very dense, transversely confluent punctures, forming small transverse striae; punctation sparser, separate and less transverse only towards the ridges of primary costae, at the humeral protuberance and behind the scutellum; epipleura rather narrow, disappearing shortly before apical curvature; marginal membrane starting near middle of metacoxa. Pygidium triangular, widely rounded at apex; somewhat tumid; with very transverse punctures which are adjacent to each other or confluate, forming a striolation which is arranged concentrically around the tumidity; each puncture bearing a microscopic seta (50x); with sparse long yellow setae ( 0.13-0.45 mm in length) in apical portion. Mesoventrite very narrow and somewhat concave between the approximated mesocoxae. Metaventrite with very dense, transversely confluent punctation; with sparse, rather short, yellowish brown setae ( 0.05-0.125 mm in length) in lateral portions; sparsely punctate and almost glabrous in the middle. Abdominal ventrites transversely punctate in the middle, densely transversely striolate at the sides; 2nd to 4th ventrite with a transverse row of semierect brownish setae ( 0.05-0.13 mm in length) in the apical half. Legs slender and long; protibia bidentate; terminal tooth somewhat prolonged and weakly obtuse at apex; protarsal segment 5 with a sharp tooth at basal 3/5; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slen- der, the lower branch broad with an obtuse angle at its ventral margin; the upper branch of outer mesotarsal claw slightly more slender than the lower one. Female : Sides of pronotum distinctly more rounded; pygidium not tumid; terminal tooth of protibia spatulate and rounded at apex; modified protarsal claw less enlarged, without ventral angle. Diagnosis : Anomala imperspicabilis spec. nov. is very similar to Anomala aulax ( WIEDEMANN, 1823 ) . There appear to be no external characters that permit the separation of these two species. Details of the aedeagus, however, is distinctly different (fig. 13): the upper apex of each of the parameres ends in a branched, thorny structure, whereas it has a laterally curved, spine-like structure in A. aulax . The lower apex of each paramere is spatulate in A. imperspicabilis but hook-shaped in A. aulax . Anomala imperspicabilis is also similar to Anomala delavayi FAIRMAIRE, 1886 , Anomala profundisulca LIN, 2002 , and Anomala aulacoides OHAUS, 1915 , but differs from these species significantly in the shape of the aedeagus. Anomala profundisulca and A. aulacoides have distinctly more convex elytral intervals than A. imperspicabilis , and A. delavayi generally has orange-brown femora and tibiae, strongly contrasting with the metallic green venter. Remarks : All specimens of A. imperspicabilis found in northern Laos have a broad yellow margin on the pronotum and elytra; however, there appear to be no differences in the aedeagus. The taxonomic value of this color difference is yet unclear. More material from other localities is required to resolve this issue. Etymology : The Latin specific epithet translates as “inscrutable” or “inapprehensible”. It is an allusion to the unclear relation between the two known populations of the new species.