The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist Author Freese, Edwin L. 33493 “ S ” Avenue Adel, Iowa 50003 Author Veal, Doug A. 168 Thompson Drive SE Cedar Rapids, Iowa 52403 Author Lago, Paul K. Biology Department, University of Mississippi University, Mississippi 38677 text Insecta Mundi 2020 2020-08-28 2020 787 1 83 journal article 10.5281/zenodo.5353792 1942-1354 5353792 88D45C5E-0E49-4F02-87A4-C4FA55A83B2F Phyllophaga knochii (Schonherr and Gyllenhal, 1817) Luginbill and Painter 1953 ; Downie and Arnett 1996 ; Harpootlian 2001 ; CASC; DAVC. Collection dates: May, July. Specimen label data: Iowa City , Johnson Co. , Iowa , 5 May 1896 , Wickham ( A. Fenyes Collection , CASSF ) ; Waubonsie State Park , Fremont Co. , Iowa , 2 July 2012 , combination light, DAV determined MJP 2012 ( DAVC ) . County records ( Figure 139 ): Fremont, Johnson.