Growth, inter- and intraspecific variation, palaeobiogeography, taphonomy and systematics of the Cenozoic ghost shrimp Glypturus Author Klompmaker, A. Florida Museum of Natural History, University of Florida, 1659 Museum Road, PO Box 117800, Gainesville, FL 32611, USA; Author Hy ̆ z, Matu ̆ s Author , Roger W. Port Author Kowalew, nd Michał text Journal of Systematic Palaeontology 2016 2015-03-16 14 2 99 126 http://dx.doi.org/10.1080/14772019.2015.1009505 journal article 10.1080/14772019.2015.1009505 1478-0941 PMC4724948 26819570 10890716 C7F0C071-F2AD-4684-B277-037B6F91BF0E Glypturus aff. G. toulai ( Rathbun, 1919 ) 2013 Glypturus toulai (Rathbun) ; Hy̆zńy et al.: 132, fig. 6. Remarks. Hy̆zńy et al. (2013) ascribed specimens from the lower Miocene Culebra Formation in Panama to G. toulai based on the extent of the tuberculation on the propodus. The limited number of specimens and their inadequate preservation does not allow rigorous bi- and multivariate analyses to reassess their placement. One possible difference with G. toulai from the upper Miocene Gatun Formation in Panama is the extent of the tubercles on the inner side of the propodus (Hy̆zńy et al. 2013, fig. 6). Also given the long interval between deposition of the Gatun and Culebra formations (~10 Ma, see Coates 1999 ; Kirby et al. 2008 ) relative to the other, younger species its ascription to G. toulai must be regarded as preliminary. Therefore, we herein propose the more conservative Glypturus aff. G. toulai until more material is studied.