A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae) Author Sabu, Thomas K. Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com sabukthomas@gmail.com Author Abhitha, Prabhakar Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com Author Zhao, Danyang text Zootaxa 2008 2008-04-09 1745 1 30 46 https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3 journal article 10.11646/zootaxa.1745.1.3 1175­5334 5108342 Helluodes westwoodi Chaudoir, 1869 ( Figs. 12–14 ) Helluodes westwoodi Chaudoir, 1869: 203 . Head. Labrum very large, as wide as long, anterior margin distinctly emarginated in the middle, with a row of 6 marginal setae. Maxillary palpomere 4 flattened, short, inflexed medially and internal surface with distinct, deep puncture; roundly expanded from basal one third, roundly narrowed at apex. Mentum with bifid median tooth. Ligula elongate, setose with four elongated apical setae, three lateral seta on either side; distinctly broader in the middle. Pronotum. Large and slightly wider than head; less densely pubescent and sparsely punctate on disc; apical angles protruded, expanded, flexed inwardly, more rounded and slightly acute on summit; anterior and posterior transverse sulci shallow; lateral margins widely explanate, distinctly sinuated anterior to posterior angles or at some distance from base; posterior angles expanded, reflexed strongly and sharp at apex; basal foveae deep and wide, stretching forward as deep lateral furrows; base narrower than apex, distinctly oblique laterally, weakly sinuated medially. Elytra. Apices obliquely truncate with wide membranous margins; outer angles sinuate; elytra with intervals sparsely punctate and pubescent, lateral edges broad. Legs. Short, front tibiae emarginated at apex. Venter. Punctate-pubescent except the glabrous posterior area of prosternum. Sternites 3 to 6 in male and female with one pair of paramedian setae. Genitalia. As for genus. Measurements. Male (n=2), TLA = 26.5 and 27.5; TLB = 24.0 and 25.0; TW = 8.5 and 10.0; PL = 3.0 and 4.0; PW = 6.0 and 7.0; EL = 14.0 and 15.0. Female (n=2), TLA = 25.0 and 26.0; TLB = 22.0 and 23.0; TW = 9.0; PL = 3.2; PW = 5.5; EL = 14.0 and 14.5. Specimens examined (n=4). Male labeled “Ex-Musaeo Chaudoir” / “westwoodii Chaud. Deccan, S. Stevens ” (designated as Lectotype in the present work); 1 female with the same label as that of Lectotype , and 1 male and 1 female labeled “ Ceylon ”. All specimens are in the Chaudoir collections of MNHN . Distribution. South India (Deccan), Sri Lanka (exact locality in both regions unknown). Remarks. Distinct from other two species with distinctly emarginate labrum; ligula broader in the middle; pronotum with rounded but slightly acute anterior angles, raised lateral margins with distinct sinuation anterior to posterior angles.