<document id="14CBCDBD71A699EA523BFCACD2D94813" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236277FFD22DEFB164FB55594B" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena dorsigera" docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="50" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="D9247131AD3349EF7DCABACD5C34E567">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="DC730AE214C5BAA9F716256BC9AC42A7">Shi, Hongliang</mods:namePart>
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<mods:namePart id="E6C4B25E4AACCFF766A8A72499C7BA64">Liang, Hongbin</mods:namePart>
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5. 
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<emphasis id="B95A1B276277FFD22DCDB164FE615E63" bold="true" box="[181,390,798,825]" italics="true" pageId="49" pageNumber="50">Parena dorsigera</emphasis>
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species group
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This species group contains two species distributed throughout 
<collectingCountry id="F33987A56277FFD22E2BB11CFC5A5EDB" box="[851,957,871,897]" name="China" pageId="49" pageNumber="50">East Asia</collectingCountry>
, north to 
<collectingCountry id="F33987A56277FFD2295BB11CFB815EDB" box="[1059,1126,871,897]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
, south to 
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and 
<collectingCountry id="F33987A56277FFD2284EB11CFA7F5EDB" box="[1334,1432,871,897]" name="Vietnam" pageId="49" pageNumber="50">Vietnam</collectingCountry>
, and west to 
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. They are characterized by: dorsum uniformly brown, elytra with indistinct middle paler patch in a few specimens; pronotum nearly quadrate; elytral apices truncate, outer apical angles distinct (
<figureCitation id="1315DBB06277FFD229B8B1D4FAD05E93" box="[1216,1335,942,969]" captionStart="FIGURE 5" captionStartId="12.[151,250,1858,1882]" captionTargetBox="[185,1402,181,1834]" captionTargetId="figure-28@12.[185,1402,181,1834]" captionTargetPageId="12" captionText="FIGURE 5. Elytral apex (left) of Parena spp. A P. (Parena) africana (Alluaud), Cameroon. B P. (Parena) ruficornis sp. n., Central Africa, paratype. C P. (Parena) valeriae Facchini, holotype. D P. (Parena) madagascariensis (Alluaud), Madagascar. E P. (Bothynoptera) tesari (Jedlička), Taiwan. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Bothynoptera) kurosai Habu, Japan. H P. (Bothynoptera) dorsigera (Schaum), Yunnan. I P. (Bothynoptera) dorsigera (Schaum), Beijing. J P. (Bothynoptera) dorsigera (Schaum), Japan. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.7958947" httpUri="https://zenodo.org/record/7958947/files/figure.png" pageId="49" pageNumber="50">Figs 5G–J</figureCitation>
); median lobe of aedeagus stout to very stout (AL/AW 3.5–4.2), apical lamella short, coniform; gonocoxite II of ovipositor rectangular, apex with five to seven ensiform setae, subequally arranged or grouped at angles.
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