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<mods:title>High cryptic diversity in the kleptoparasitic genus Hyalorisia Dall, 1889 (Littorinimorpha: Capulidae) with the description of nine new species from the Indo-West Pacific</mods:title>
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<mods:namePart>Fassio, Giulia</mods:namePart>
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<mods:namePart>Russini, Valeria</mods:namePart>
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<mods:namePart>Modica, Maria Vittoria</mods:namePart>
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<mods:title>Rijksuniversiteit te Gent. Fakulteit van de Landbouwkundige en Toegepaste Biologische Wetenschappen</mods:title>
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<taxonomicName authority="(Dall, 1889)" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[1000,1319,1125,1147]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="galea">
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<emphasis bold="true" box="[1000,1182,1125,1147]" italics="true" pageId="10" pageNumber="411">Hyalorisia galea</emphasis>
(
<bibRefCitation author="DALL, W. H." box="[1197,1311,1125,1146]" editor="C. D. Sigsbee &amp; J. R. Bartlet" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="10" pageNumber="411" pagination="1 - 492" part="18" refId="ref14958" refString="DALL, W. H. 1889. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlet. Bulletin of the Museum of Comparative Zoology at Harvard College, 18: 1 - 492, pls 10 - 40." title="Reports on the results of dredging, under the supervision of Alexander Agassiz" type="journal article" volumeTitle="the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander" year="1889">Dall, 1889</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection box="[982,1338,1152,1174]" pageId="10" pageNumber="411" type="description">
<paragraph blockId="10.[982,1338,1125,1174]" box="[982,1338,1152,1174]" pageId="10" pageNumber="411">
<heading box="[982,1338,1152,1174]" centered="true" fontSize="9" level="5" pageId="10" pageNumber="411" reason="4">
MOTU-1 (
<figureCitation box="[1090,1164,1152,1174]" captionStart="Figure 3" captionStartId="5.[154,219,1780,1799]" captionTargetBox="[173,1458,147,1753]" captionTargetId="figure-4@5.[173,1458,147,1755]" captionTargetPageId="5" captionText="Figure 3. Protoconchs of Hyalorisia. A. Hyalorisia galea, MNHN-IM-2013-61536. B. Hyalorisia neocaledonica n. sp., MNHN-IM-2013-65907. C. Hyalorisia tosaensis, MNHN-IM-2007-34794. D. Hyalorisia nupta n. sp., MNHN-IM-2007-34005. E. Hyalorisia lehibe n. sp., MNHN-IM-2007-39212. F. Hyalorisia nanhaiensis n. sp., MNHN-IM-2013-34796. G. Hyalorisia kely n. sp., MNHN-IM-2007-38809. H. Hyalorisia melanesica n. sp., MNHN-IM-2013-58359. I. Hyalorisia profunda n. sp., MNHN-IM-2007-33828. J. Hyalorisia madagascarensis n. sp., MNHN-IM-39292. K. Hyalorisia solomonensis n. sp., MNHN-IM-2007-34215. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4464906" httpUri="https://zenodo.org/record/4464906/files/figure.png" pageId="10" pageNumber="411">Figs 3A</figureCitation>
, 
<figureCitation box="[1174,1204,1152,1173]" captionStart="Figure 4" captionStartId="6.[156,221,1777,1796]" captionTargetBox="[175,1459,148,1752]" captionTargetId="figure-4@6.[174,1459,148,1752]" captionTargetPageId="6" captionText="Figure 4. Shells of Hyalorisia. A. Hyalorisia galea, MNHN-IM-2013-60195, N of Grande-Terre, Guadeloupe (L = 13 mm). B. Hyalorisia neocaledonica n. sp., holotype, MNHN-IM-2013-65645, Lord Howe Rise, New Caledonia (L = 9.5 mm). C. Hyalorisia tosaensis, MNHN-IM-2013-58352, New Ireland, Papua New Guinea (L = 15 mm). D. Hyalorisia nupta n. sp., holotype, MNHN-IM-2007-34010, Bellona Reefs, New Caledonia (L = 18.5 mm). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4464908" httpUri="https://zenodo.org/record/4464908/files/figure.png" pageId="10" pageNumber="411">4A</figureCitation>
, 
<figureCitation box="[1214,1263,1152,1173]" captionStart="Figure 7" captionStartId="9.[154,219,1780,1799]" captionTargetBox="[173,1458,148,1755]" captionTargetId="figure-4@9.[173,1458,148,1755]" captionTargetPageId="9" captionText="Figure 7. Radulae of Hyalorisia (A–I) and detail of the rachidian tooth (J–R). A, J. Hyalorisia galea MNHN-IM-2013-60195. B, K. Hyalorisia neocaledonica n. sp., MNHN-IM-2013-65647. C, L. Hyalorisia tosaensis, MNHN-IM-2013-58352. D, M. Hyalorisia nupta n. sp., MNHN-IM-2013-68952. E, N. Hyalorisia nanhaiensis n. sp., MNHN-IM-2013-59697. F, O. Hyalorisia kely n. sp., MNHN-IM-2007-38809. G, P. Hyalorisia melanesica n. sp., MNHN-IM-2007-34205. H, Q. Hyalorisia madagascarensis n. sp., MNHN-IM-2007-39090. I, R. Hyalorisia solomonensis n. sp., MNHN-IM-2007-34215. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4464916" httpUri="https://zenodo.org/record/4464916/files/figure.png" pageId="10" pageNumber="411">7A, J</figureCitation>
, 
<figureCitation box="[1273,1331,1152,1174]" captionStart="Figure 8" captionStartId="11.[154,219,1761,1780]" captionTargetBox="[172,1460,146,1736]" captionTargetId="figure-4@11.[172,1460,146,1736]" captionTargetPageId="11" captionText="Figure 8. A–C. Holotype of Capulus (Hyalorisia) galea Dall, 1889, USNM 508724, off Barbados. D–F. Holotype of Capulus (Hyalorisia) tosaensis Otuka, 1939, Seto Marine Biological Laboratory, collected from Bay of Tosa, Japan. G–I. Syntype of Capulus fragilis E. A. Smith, 1904, NHMUK 1904.6.15.136, Laccadive Sea. J. Original figure of Capulus fallax S. V. Wood, 1842 (Wood, 1842: pl. 17, fig. 4a, b), Lower Pliocene of England. K–N. Holotype of Capulus (Hyalorisia) nettlesi J. E. Robinson, 1983, PRI 30058, Upper Eocene of Mississippi. Scale bars: A–E, G–I, K–N = 10 mm; F = 5 mm. Photo credits: A–C, USNM; D–F, Kyoto University (taken by Ryutaro Goto); G–I, NHMUK (taken by Harry Taylor); K–N, Paleontological Research Institution, Ithaca, NY." figureDoi="http://doi.org/10.5281/zenodo.4464918" httpUri="https://zenodo.org/record/4464918/files/figure.png" pageId="10" pageNumber="411">8C, D</figureCitation>
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<treatmentCitation author="DALL, W. H." box="[841,1183,1209,1228]" page="288" pageId="10" pageNumber="411" year="1889">
<taxonomicName authority="Dall, 1889: 288" authorityName="Dall" authorityPageNumber="288" authorityYear="1889" box="[841,1183,1209,1228]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="galea" subGenus="Hyalorisia">
<emphasis box="[841,902,1209,1228]" italics="true" pageId="10" pageNumber="411">Capulus</emphasis>
(
<emphasis box="[913,990,1209,1228]" italics="true" pageId="10" pageNumber="411">Hyalorisia</emphasis>
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<emphasis box="[1002,1039,1209,1228]" italics="true" pageId="10" pageNumber="411">galea</emphasis>
<bibRefCitation author="DALL, W. H." box="[1046,1183,1209,1228]" editor="C. D. Sigsbee &amp; J. R. Bartlet" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="10" pageNumber="411" pagination="1 - 492" part="18" refId="ref14958" refString="DALL, W. H. 1889. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlet. Bulletin of the Museum of Comparative Zoology at Harvard College, 18: 1 - 492, pls 10 - 40." title="Reports on the results of dredging, under the supervision of Alexander Agassiz" type="journal article" volumeTitle="the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander" year="1889">Dall, 1889: 288</bibRefCitation>
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, pl. 14, fig. 3 [
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off 
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<emphasis box="[841,964,1257,1278]" italics="true" pageId="10" pageNumber="411">Capulus galeus</emphasis>
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(
<emphasis box="[976,997,1258,1279]" italics="true" pageId="10" pageNumber="411">sic</emphasis>
)— 
<treatmentCitation author="ROBINSON, J. E." box="[1027,1204,1257,1279]" page="9" pageId="10" pageNumber="411" year="1983">
<bibRefCitation author="ROBINSON, J. E." box="[1027,1204,1257,1279]" journalOrPublisher="Mississippi Geology" pageId="10" pageNumber="411" pagination="9 - 12" part="3" refId="ref16300" refString="ROBINSON, J. E. 1983. New and interesting Eocene molluscan species from the Moodys Branch Formation. Mississippi Geology, 3: 9 - 12." title="New and interesting Eocene molluscan species from the Moodys Branch Formation" type="journal article" year="1983">Robinson, 1983: 9</bibRefCitation>
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<treatmentCitationGroup pageId="10" pageNumber="411">
<taxonomicName authority="(Dall, 1889)" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[841,1070,1282,1301]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="galea">
<emphasis box="[841,962,1282,1301]" italics="true" pageId="10" pageNumber="411">Hyalorisia galea</emphasis>
(
<bibRefCitation author="DALL, W. H." box="[974,1066,1282,1301]" editor="C. D. Sigsbee &amp; J. R. Bartlet" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="10" pageNumber="411" pagination="1 - 492" part="18" refId="ref14958" refString="DALL, W. H. 1889. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlet. Bulletin of the Museum of Comparative Zoology at Harvard College, 18: 1 - 492, pls 10 - 40." title="Reports on the results of dredging, under the supervision of Alexander Agassiz" type="journal article" volumeTitle="the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander" year="1889">Dall, 1889</bibRefCitation>
)
</taxonomicName>
— 
<treatmentCitation author="GRACIA CLAVIJO, M. A. &amp; ARDILA, N. E. &amp; DIAZ, J. M." box="[1092,1443,1282,1303]" page="52" pageId="10" pageNumber="411" year="2004">
<bibRefCitation author="GRACIA CLAVIJO, M. A. &amp; ARDILA, N. E. &amp; DIAZ, J. M." box="[1092,1443,1282,1303]" journalOrPublisher="Iberus" pageId="10" pageNumber="411" pagination="43 - 75" part="22" refId="ref15386" refString="GRACIA CLAVIJO, M. A., ARDILA, N. E. &amp; DIAZ, J. M. 2004. Gastropods collected along the continental slope of the Colombian Caribbean during the INVEMAR-Macrofauna campaigns (1998 - 2001). Iberus, 22: 43 - 75." title="Gastropods collected along the continental slope of the Colombian Caribbean during the INVEMAR-Macrofauna campaigns (1998 - 2001)" type="journal article" year="2004">Gracia Clavijo, Ardila &amp; Díaz, 2004: 52</bibRefCitation>
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, fig. 22. 
<treatmentCitation author="ROSENBERG, G. &amp; MORETZSOHN, F. &amp; GARCIA, E. F." box="[906,1298,1306,1326]" page="635" pageId="10" pageNumber="411" year="2009">
<bibRefCitation author="ROSENBERG, G. &amp; MORETZSOHN, F. &amp; GARCIA, E. F." box="[906,1298,1306,1326]" editor="D. L. Felder &amp; D. K. Camp" journalOrPublisher="Texas A &amp; M University Press, College Station, TX" pageId="10" pageNumber="411" pagination="579 - 699" refId="ref16438" refString="ROSENBERG, G., MORETZSOHN, F. &amp; GARCIA, E. F. 2009. Gastropoda (Mollusca) of the Gulf of Mexico. In: Gulf of Mexico-origins, waters, and biota. Vol. 1: Biodiversity (D. L. Felder &amp; D. K. Camp, eds), pp. 579 - 699. Texas A &amp; M University Press, College Station, TX." title="Gastropoda (Mollusca) of the Gulf of Mexico" type="book chapter" volumeTitle="Gulf of Mexico-origins, waters, and biota. Vol. 1: Biodiversity" year="2009">Rosenberg, Moretzsohn &amp; García, 2009: 635</bibRefCitation>
</treatmentCitation>
. 
<treatmentCitation author="DACCARETT, E. Y. &amp; BOSSIO, V. M. S." page="64" pageId="10" pageNumber="411" year="2011">
<bibRefCitation author="DACCARETT, E. Y. &amp; BOSSIO, V. M. S." journalOrPublisher="L'Informatore Piceno, Ancona" pageId="10" pageNumber="411" refId="ref14929" refString="DACCARETT, E. Y. &amp; BOSSIO, V. M. S. 2011. Colombian seashells from the Caribbean Sea. L'Informatore Piceno, Ancona." title="Colombian seashells from the Caribbean Sea" type="book" year="2011">Daccarett &amp; Bossio, 2011: 64</bibRefCitation>
</treatmentCitation>
. 
<treatmentCitation author="GARCIA, E. F." box="[958,1103,1330,1349]" page="34" pageId="10" pageNumber="411" year="2012">
<bibRefCitation author="GARCIA, E. F." box="[958,1103,1330,1349]" journalOrPublisher="American Conchologist" pageId="10" pageNumber="411" pagination="34 - 37" part="40" refId="ref15351" refString="GARCIA, E. F. 2012. Noteworthy offshore mollusks from the north-central Gulf of Mexico, including geographical extensions and a generic reassignment. American Conchologist, 40: 34 - 37." title="Noteworthy offshore mollusks from the north-central Gulf of Mexico, including geographical extensions and a generic reassignment" type="journal article" year="2012">García, 2012: 34</bibRefCitation>
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, fig. 4.
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</paragraph>
<paragraph blockId="10.[841,1480,1209,1376]" box="[841,1226,1354,1376]" pageId="10" pageNumber="411">
<treatmentCitationGroup box="[841,1226,1354,1376]" pageId="10" pageNumber="411">
<taxonomicName box="[841,992,1354,1375]" class="Gastropoda" family="Phenacolepadidae" genus="Plesiothyreus" kingdom="Animalia" order="Cycloneritimorpha" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="galea">
<emphasis box="[841,992,1354,1375]" italics="true" pageId="10" pageNumber="411">Plesiothyreus galea</emphasis>
</taxonomicName>
— 
<treatmentCitation author="CHRISTIAENS, J." box="[1016,1222,1354,1376]" page="22" pageId="10" pageNumber="411" year="1989">
<bibRefCitation author="CHRISTIAENS, J." box="[1016,1222,1354,1376]" journalOrPublisher="Gloria Maris" pageId="10" pageNumber="411" pagination="17 - 48" part="28" refId="ref14863" refString="CHRISTIAENS, J. 1989. The Phenacolepadidae. Gastropoda: Neritoidea, part 2. Gloria Maris, 28: 17 - 48." title="The Phenacolepadidae. Gastropoda: Neritoidea, part 2" type="journal article" year="1989">Christiaens, 1989: 22</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="411" type="materials_examined">
<paragraph blockId="10.[841,1480,1397,1791]" pageId="10" pageNumber="411">
<emphasis box="[841,997,1397,1418]" italics="true" pageId="10" pageNumber="411">Material examined</emphasis>
: 
<materialsCitation ID-GBIF-Occurrence="3025459310" collectionCode="MNHN" pageId="10" pageNumber="411" specimenCount="2" typeStatus="holotype">
The 
<typeStatus box="[1060,1143,1397,1419]" pageId="10" pageNumber="411">holotype</typeStatus>
(only photographic images examined; 
<figureCitation box="[901,961,1423,1445]" captionStart="Figure 8" captionStartId="11.[154,219,1761,1780]" captionTargetBox="[172,1460,146,1736]" captionTargetId="figure-4@11.[172,1460,146,1736]" captionTargetPageId="11" captionText="Figure 8. A–C. Holotype of Capulus (Hyalorisia) galea Dall, 1889, USNM 508724, off Barbados. D–F. Holotype of Capulus (Hyalorisia) tosaensis Otuka, 1939, Seto Marine Biological Laboratory, collected from Bay of Tosa, Japan. G–I. Syntype of Capulus fragilis E. A. Smith, 1904, NHMUK 1904.6.15.136, Laccadive Sea. J. Original figure of Capulus fallax S. V. Wood, 1842 (Wood, 1842: pl. 17, fig. 4a, b), Lower Pliocene of England. K–N. Holotype of Capulus (Hyalorisia) nettlesi J. E. Robinson, 1983, PRI 30058, Upper Eocene of Mississippi. Scale bars: A–E, G–I, K–N = 10 mm; F = 5 mm. Photo credits: A–C, USNM; D–F, Kyoto University (taken by Ryutaro Goto); G–I, NHMUK (taken by Harry Taylor); K–N, Paleontological Research Institution, Ithaca, NY." figureDoi="http://doi.org/10.5281/zenodo.4464918" httpUri="https://zenodo.org/record/4464918/files/figure.png" pageId="10" pageNumber="411">Fig. 8</figureCitation>
A–C)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3025459301" collectionCode="MNHN" pageId="10" pageNumber="411" specimenCode="MNHN-IM-2013-60195, MNHN-IM-2013-61224, MNHN-IM-2013-61525, MNHN-IM-2013-61536" specimenCount="1" typeStatus="other material">
and other material (
<specimenCode box="[1236,1475,1423,1445]" pageId="10" pageNumber="411">MNHN-IM-2013-60195</specimenCode>
, 
<specimenCode box="[841,1081,1450,1472]" pageId="10" pageNumber="411">MNHN-IM-2013-61224</specimenCode>
, 
<specimenCode box="[1096,1335,1450,1472]" pageId="10" pageNumber="411">MNHN-IM-2013-61525</specimenCode>
and 
<specimenCode pageId="10" pageNumber="411">MNHN-IM-2013-61536</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="414" pageId="10" pageNumber="411" type="diagnosis">
<paragraph blockId="10.[841,1480,1397,1791]" pageId="10" pageNumber="411">
<emphasis box="[841,1018,1503,1524]" italics="true" pageId="10" pageNumber="411">COI-based diagnosis</emphasis>
: Ten diagnostic (autapomorphic) positions in alignment: 74(C), 145(T), 184(G), 208(G), 304(T), 418(A), 424(T), 601(C), 622(A) and 637(T).
</paragraph>
<paragraph blockId="10.[841,1480,1397,1791]" pageId="10" pageNumber="411">
<emphasis box="[841,1256,1583,1604]" italics="true" pageId="10" pageNumber="411">
Shell (data for 
<typeStatus box="[968,1033,1583,1604]" pageId="10" pageNumber="411">holotype</typeStatus>
shown within parentheses)
</emphasis>
: Size medium to large for the genus: length 13–25 (15.5) mm, width 9–21 (10.7) mm, height 2.5–8 (4.9) mm; thin, rounded, ellipsoidal or ovoidal, depressed; white shell surface with smooth yellowish periostracum; anterior margin rounded or very slightly angled; protoconch of 1– 1.2 whorls, diameter of nucleus 
<quantity box="[1146,1289,1716,1737]" metricMagnitude="-4" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="10" pageNumber="411" unit="mm" value="0.185" valueMax="0.2" valueMin="0.17">0.17–0.20 mm</quantity>
; teleoconch smooth except for distinct commarginal growth ridges; apertural margin thin, interior glossy white; lamella narrow, width 0.14–2 (0.14) mm.
</paragraph>
<paragraph blockId="10.[841,1479,1822,1871]" pageId="10" pageNumber="411">
<emphasis box="[841,1091,1822,1844]" italics="true" pageId="10" pageNumber="411">Soft parts (ethanol preserved)</emphasis>
: Head, cephalic tentacles, pseudoproboscis and mantle whitish.
</paragraph>
<paragraph blockId="10.[841,1479,1902,1950]" lastBlockId="13.[154,792,147,196]" lastPageId="13" lastPageNumber="414" pageId="10" pageNumber="411">
<emphasis box="[841,902,1902,1923]" italics="true" pageId="10" pageNumber="411">Radula</emphasis>
: Taenioglossate, 2-1-R-1-2; marginal teeth sickle shaped; lateral teeth triangular, outer edge smooth or with few denticles, inneedge smooth or with one denticle; rachidian tooth triangular, highly denticulated on both edges.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4464918" ID-Zenodo-Dep="4464918" httpUri="https://zenodo.org/record/4464918/files/figure.png" pageId="11" pageNumber="412" startId="11.[154,219,1761,1780]" targetBox="[172,1460,146,1736]" targetPageId="11">
<paragraph blockId="11.[154,1477,1761,1878]" pageId="11" pageNumber="412">
<emphasis bold="true" box="[154,297,1761,1781]" pageId="11" pageNumber="412">Figure 8. A–C.</emphasis>
Holotype of 
<taxonomicName authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[410,703,1762,1781]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="galea" subGenus="Hyalorisia">
<emphasis box="[410,471,1762,1781]" italics="true" pageId="11" pageNumber="412">Capulus</emphasis>
(
<emphasis box="[482,559,1762,1781]" italics="true" pageId="11" pageNumber="412">Hyalorisia</emphasis>
) 
<emphasis box="[570,607,1762,1781]" italics="true" pageId="11" pageNumber="412">galea</emphasis>
<bibRefCitation author="DALL, W. H." box="[613,703,1762,1781]" editor="C. D. Sigsbee &amp; J. R. Bartlet" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="11" pageNumber="412" pagination="1 - 492" part="18" refId="ref14958" refString="DALL, W. H. 1889. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlet. Bulletin of the Museum of Comparative Zoology at Harvard College, 18: 1 - 492, pls 10 - 40." title="Reports on the results of dredging, under the supervision of Alexander Agassiz" type="journal article" volumeTitle="the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander" year="1889">Dall, 1889</bibRefCitation>
</taxonomicName>
, USNM 508724, off Barbados. 
<emphasis bold="true" box="[978,1022,1762,1781]" pageId="11" pageNumber="412">D–F.</emphasis>
Holotype of 
<taxonomicName authority="Otuka, 1939" authorityName="Otuka" authorityYear="1939" box="[1136,1472,1762,1781]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="tosaensis" subGenus="Hyalorisia">
<emphasis box="[1136,1197,1762,1781]" italics="true" pageId="11" pageNumber="412">Capulus</emphasis>
(
<emphasis box="[1208,1285,1762,1781]" italics="true" pageId="11" pageNumber="412">Hyalorisia</emphasis>
) 
<emphasis box="[1296,1358,1762,1780]" italics="true" pageId="11" pageNumber="412">tosaensis</emphasis>
<bibRefCitation author="OTUKA, Y." box="[1364,1472,1762,1781]" journalOrPublisher="Venus" pageId="11" pageNumber="412" pagination="89 - 98" part="9" refId="ref15933" refString="OTUKA, Y. 1939. On some non-sculptured species of the genus Capulus. Venus, 9: 89 - 98." title="On some non-sculptured species of the genus Capulus" type="journal article" year="1939">Otuka, 1939</bibRefCitation>
</taxonomicName>
, Seto Marine Biological Laboratory, collected from Bay of Tosa, Japan. 
<emphasis bold="true" box="[748,791,1785,1804]" pageId="11" pageNumber="412">G–I.</emphasis>
Syntype of 
<taxonomicName authority="E. A. Smith, 1904" authorityName="E. A. Smith" authorityYear="1904" box="[889,1160,1786,1805]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="fragilis">
<emphasis box="[889,1004,1786,1805]" italics="true" pageId="11" pageNumber="412">Capulus fragilis</emphasis>
E. A. 
<bibRefCitation author="SMITH, E. A." box="[1056,1160,1786,1805]" journalOrPublisher="On Mollusca from the Bay of Bengal and the Arabian Sea. Annals and Magazine of Natural History, Including Zoology, Botany, and Geology" pageId="11" pageNumber="412" pagination="245 - 264" part="18" refId="ref16838" refString="SMITH, E. A. 1904. Natural history notes from R. I. M. S. ' Investigator. ' - Series III, No. 10. On Mollusca from the Bay of Bengal and the Arabian Sea. Annals and Magazine of Natural History, Including Zoology, Botany, and Geology, Series 7, 18: 245 - 264." title="Natural history notes from R. I. M. S. ' Investigator" type="journal article" year="1904">Smith, 1904</bibRefCitation>
</taxonomicName>
, NHMUK 1904.6.15.136, Laccadive Sea. 
<emphasis bold="true" box="[196,211,1810,1829]" pageId="11" pageNumber="412">J.</emphasis>
Original figure of 
<taxonomicName authority="S. V. Wood, 1842" authorityName="S. V. Wood" authorityYear="1842" box="[373,637,1809,1829]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="fallax">
<emphasis box="[373,481,1809,1828]" italics="true" pageId="11" pageNumber="412">Capulus fallax</emphasis>
S. V. Wood, 1842
</taxonomicName>
(Wood, 1842: pl. 17, fig. 4a, b), Lower Pliocene of England. 
<emphasis bold="true" box="[1159,1209,1810,1829]" pageId="11" pageNumber="412">K–N.</emphasis>
Holotype of 
<taxonomicName authority="J. E. Robinson, 1983" authorityName="J. E. Robinson" authorityYear="1983" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="nettlesi" subGenus="Hyalorisia">
<emphasis box="[1323,1384,1809,1828]" italics="true" pageId="11" pageNumber="412">Capulus</emphasis>
(
<emphasis box="[1395,1472,1809,1828]" italics="true" pageId="11" pageNumber="412">Hyalorisia</emphasis>
) 
<emphasis box="[154,203,1833,1852]" italics="true" pageId="11" pageNumber="412">nettlesi</emphasis>
J. E. 
<bibRefCitation author="ROBINSON, J. E." box="[249,384,1833,1853]" journalOrPublisher="Mississippi Geology" pageId="11" pageNumber="412" pagination="9 - 12" part="3" refId="ref16300" refString="ROBINSON, J. E. 1983. New and interesting Eocene molluscan species from the Moodys Branch Formation. Mississippi Geology, 3: 9 - 12." title="New and interesting Eocene molluscan species from the Moodys Branch Formation" type="journal article" year="1983">Robinson, 1983</bibRefCitation>
</taxonomicName>
, PRI 30058, Upper Eocene of Mississippi. Scale bars: 
<emphasis bold="true" box="[848,886,1833,1853]" pageId="11" pageNumber="412">A–E</emphasis>
, 
<emphasis bold="true" box="[897,933,1833,1853]" pageId="11" pageNumber="412">G–I</emphasis>
, 
<emphasis bold="true" box="[943,987,1834,1853]" pageId="11" pageNumber="412">K–N</emphasis>
= 10 mm; 
<emphasis bold="true" box="[1085,1097,1834,1853]" pageId="11" pageNumber="412">F</emphasis>
= 5 mm. Photo credits: 
<emphasis bold="true" box="[1307,1347,1833,1853]" pageId="11" pageNumber="412">A–C</emphasis>
, USNM; 
<emphasis bold="true" box="[1432,1472,1834,1853]" pageId="11" pageNumber="412">D–F</emphasis>
, Kyoto University (taken by Ryutaro Goto); 
<emphasis bold="true" box="[521,558,1857,1877]" pageId="11" pageNumber="412">G–I</emphasis>
, NHMUK (taken by Harry Taylor); 
<emphasis bold="true" box="[871,915,1857,1877]" pageId="11" pageNumber="412">K–N</emphasis>
, Paleontological Research Institution, Ithaca, NY.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4464920" ID-Zenodo-Dep="4464920" httpUri="https://zenodo.org/record/4464920/files/figure.png" pageId="12" pageNumber="413" startId="12.[156,221,1777,1796]" targetBox="[174,1459,146,1752]" targetPageId="12">
<paragraph blockId="12.[156,1479,1777,1870]" pageId="12" pageNumber="413">
<emphasis bold="true" box="[156,245,1777,1797]" pageId="12" pageNumber="413">Figure 9.</emphasis>
Bivalve hosts (
<taxonomicName authorityName="de Gregorio" authorityYear="1884" box="[371,484,1778,1796]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="12" pageNumber="413" phylum="Mollusca" rank="genus">
<emphasis box="[371,484,1778,1796]" italics="true" pageId="12" pageNumber="413">Propeamussium</emphasis>
</taxonomicName>
spp.) with foot mark (black arrows) and notch (white arrows) of 
<taxonomicName authorityName="Dall" authorityYear="1889" box="[1027,1104,1778,1797]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="413" phylum="Mollusca" rank="genus">
<emphasis box="[1027,1104,1778,1797]" italics="true" pageId="12" pageNumber="413">Hyalorisia</emphasis>
</taxonomicName>
. 
<emphasis bold="true" box="[1115,1135,1777,1796]" pageId="12" pageNumber="413">A.</emphasis>
<taxonomicName baseAuthorityName="Dautzenberg &amp; Bavay" baseAuthorityYear="1904" box="[1141,1310,1778,1797]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="sibogai">
<emphasis box="[1141,1310,1778,1797]" italics="true" pageId="12" pageNumber="413">Propeamussium sibogai</emphasis>
</taxonomicName>
, MNHN-IM-2007- 33954 (
<emphasis box="[219,231,1802,1820]" italics="true" pageId="12" pageNumber="413">L</emphasis>
= 46.7 mm), host of 
<taxonomicName authority="Fassio &amp; Russini &amp; Buge &amp; Schiaparelli &amp; Modica &amp; Bouchet &amp; Oliverio, 2020" authorityName="Fassio &amp; Russini &amp; Buge &amp; Schiaparelli &amp; Modica &amp; Bouchet &amp; Oliverio" authorityYear="2020" box="[413,480,1802,1820]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="nupta" status="n. sp.">
<emphasis box="[413,480,1802,1820]" italics="true" pageId="12" pageNumber="413">H. nupta</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[485,531,1802,1821]" pageId="12" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
(MNHN-IM-2007-34010). 
<emphasis bold="true" box="[768,790,1802,1821]" pageId="12" pageNumber="413">B.</emphasis>
<taxonomicName baseAuthorityName="E. A. Smith" baseAuthorityYear="1885" box="[795,976,1802,1821]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="caducum">
<emphasis box="[795,976,1802,1821]" italics="true" pageId="12" pageNumber="413">Propeamussium caducum</emphasis>
</taxonomicName>
, MNHN-IM-2007-39095 (
<emphasis box="[1209,1221,1802,1820]" italics="true" pageId="12" pageNumber="413">L</emphasis>
= 17.8 mm), host of 
<taxonomicName authority="Fassio &amp; Russini &amp; Buge &amp; Schiaparelli &amp; Modica &amp; Bouchet &amp; Oliverio, 2020" authorityName="Fassio &amp; Russini &amp; Buge &amp; Schiaparelli &amp; Modica &amp; Bouchet &amp; Oliverio" authorityYear="2020" box="[1402,1457,1802,1821]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="kely" status="n. sp.">
<emphasis box="[1402,1457,1802,1821]" italics="true" pageId="12" pageNumber="413">H. kely</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
(MNHN-IM-2007-39091). 
<emphasis bold="true" box="[417,469,1825,1845]" pageId="12" pageNumber="413">C, D.</emphasis>
<taxonomicName baseAuthorityName="E. A. Smith" baseAuthorityYear="1885" box="[474,626,1825,1844]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="dalli">
<emphasis box="[474,626,1825,1844]" italics="true" pageId="12" pageNumber="413">Propeamussium dalli</emphasis>
</taxonomicName>
, MNHN-IM-2013-60196 (
<emphasis box="[859,871,1826,1844]" italics="true" pageId="12" pageNumber="413">L</emphasis>
= 83 mm) with 
<taxonomicName authorityName="Dall" authorityYear="1889" box="[1013,1077,1825,1844]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="413" phylum="Mollusca" rank="species" species="galea">
<emphasis box="[1013,1077,1825,1844]" italics="true" pageId="12" pageNumber="413">H. galea</emphasis>
</taxonomicName>
(MNHN-IM-2013-60195) still attached. Photo credits: MNHN; taken by Yves Terryn (
<emphasis bold="true" box="[491,504,1849,1868]" pageId="12" pageNumber="413">A</emphasis>
, 
<emphasis bold="true" box="[515,530,1850,1869]" pageId="12" pageNumber="413">B</emphasis>
) and Laurent Charles (
<emphasis bold="true" box="[727,743,1849,1868]" pageId="12" pageNumber="413">C</emphasis>
). Scale bars = 5 mm.
</paragraph>
</caption>
<subSubSection pageId="13" pageNumber="414" type="biology_ecology">
<paragraph blockId="13.[154,792,227,462]" pageId="13" pageNumber="414">
<emphasis box="[154,194,228,249]" italics="true" pageId="13" pageNumber="414">Host</emphasis>
: Two individuals of 
<taxonomicName box="[406,491,227,249]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="galaea">
<emphasis box="[406,491,227,249]" italics="true" pageId="13" pageNumber="414">H. galaea</emphasis>
</taxonomicName>
, MNHN-IM-2013-60195 and MNHN-IM-2013-61224, were attached to two examples of 
<taxonomicName baseAuthorityName="E. A. Smith" baseAuthorityYear="1885" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="dalli">
<emphasis italics="true" pageId="13" pageNumber="414">Propeamussium dalli</emphasis>
</taxonomicName>
(MNHN-IM-2013-60196 and MNHN-IM-2013-61223, respectively). Two other specimens of 
<taxonomicName box="[608,692,307,328]" class="Gastropoda" family="Capulidae" genus="Hyalorisia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="galaea">
<emphasis box="[608,692,307,328]" italics="true" pageId="13" pageNumber="414">H. galaea</emphasis>
</taxonomicName>
(
<collectionCode box="[708,792,307,328]" pageId="13" pageNumber="414">MNHN-IM-</collectionCode>
2013-61525 and MNHN-IM-2013-61536) were collected at stations where the only 
<taxonomicName authorityName="Abbott" authorityYear="1954" box="[394,555,360,382]" class="Bivalvia" family="Propeamussiidae" kingdom="Animalia" order="Pectinoida" pageId="13" pageNumber="414" phylum="Mollusca" rank="family">Propeamussiidae</taxonomicName>
found were 
<taxonomicName baseAuthorityName="E. A. Smith" baseAuthorityYear="1885" box="[685,746,360,382]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="dalli">
<emphasis box="[685,746,360,382]" italics="true" pageId="13" pageNumber="414">P. dalli</emphasis>
</taxonomicName>
and 
<taxonomicName authority="(E. A. Smith, 1885)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1885" box="[154,583,386,409]" class="Bivalvia" family="Propeamussiidae" genus="Parvamussium" kingdom="Animalia" order="Pectinoida" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="cancellatum">
<emphasis box="[154,378,386,408]" italics="true" pageId="13" pageNumber="414">Parvamussium cancellatum</emphasis>
(E. A. Smith, 1885)
</taxonomicName>
. 
<bibRefCitation author="GARCIA, E. F." box="[595,722,387,409]" journalOrPublisher="American Conchologist" pageId="13" pageNumber="414" pagination="34 - 37" part="40" refId="ref15351" refString="GARCIA, E. F. 2012. Noteworthy offshore mollusks from the north-central Gulf of Mexico, including geographical extensions and a generic reassignment. American Conchologist, 40: 34 - 37." title="Noteworthy offshore mollusks from the north-central Gulf of Mexico, including geographical extensions and a generic reassignment" type="journal article" year="2012">García (2012</bibRefCitation>
: fig. 4) reported 
<taxonomicName authorityName="Dall" authorityYear="1889" box="[241,311,413,435]" class="Gastropoda" family="Capulidae" genus="Capulus" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="galea">
<emphasis box="[241,311,413,435]" italics="true" pageId="13" pageNumber="414">H. galea</emphasis>
</taxonomicName>
attached to a living 
<taxonomicName authorityName="de Gregorio" authorityYear="1884" box="[500,627,414,435]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="13" pageNumber="414" phylum="Mollusca" rank="genus">
<emphasis box="[500,627,414,435]" italics="true" pageId="13" pageNumber="414">Propeamussium</emphasis>
</taxonomicName>
sp. from the 
<collectingRegion box="[749,792,413,435]" country="Papua New Guinea" name="Gulf" pageId="13" pageNumber="414">Gulf</collectingRegion>
of 
<collectingRegion box="[179,253,440,462]" country="Mexico" name="Mexico" pageId="13" pageNumber="414">Mexico</collectingRegion>
, off 
<collectingRegion box="[293,390,440,462]" country="United States of America" name="Louisiana" pageId="13" pageNumber="414">Louisiana</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="414" type="distribution">
<paragraph blockId="13.[154,792,493,541]" pageId="13" pageNumber="414">
<emphasis box="[154,252,493,514]" italics="true" pageId="13" pageNumber="414">Distribution</emphasis>
: From the Louisiana Slope to 
<collectingCountry box="[603,694,493,515]" name="Barbados" pageId="13" pageNumber="414">Barbados</collectingCountry>
and the Caribbean coast of 
<collectingCountry box="[341,439,519,541]" name="Colombia" pageId="13" pageNumber="414">Colombia</collectingCountry>
, at depths of 
<quantity box="[566,678,519,541]" metricMagnitude="2" metricUnit="m" metricValue="6.16" metricValueMax="8.35" metricValueMin="3.97" pageId="13" pageNumber="414" unit="m" value="616.0" valueMax="835.0" valueMin="397.0">
397– 
<specimenCount box="[615,682,519,541]" pageId="13" pageNumber="414" type="generic">835 m.</specimenCount>
</quantity>
</paragraph>
</subSubSection>
</treatment>
</document>