Revision of the family Asteroseriidae (Anthozoa, Scleractinia, Cretaceous)
Author
Löser, Hannes
text
Geodiversitas
2013
2013-03-29
35
1
21
29
http://dx.doi.org/10.5252/g2013n1a2
journal article
6414
10.5252/g2013n1a2
c36e980f-7d7d-4b7d-a253-d3d4dc73de11
1638-9395
4537923
Genus
Actinoseris
d’Orbigny, 1849
TYPE
SPECIES
. —
Actinoseris cenomaniensis
d’Orbigny, 1849
, by monotypy. The
type
species is valid as combined description of genus and species.
RANGE
. — Hauterivian to Cenomanian.
DIAGNOSIS. — Solitary patellate corallum. Calicular outline circular.
Septa
compact. Microstructure of septa unknown.
Septa
in cross section externally slightly thicker, and slightly thinner towards the centre of the calice.Septal maximum thickness
250 µm
. Symmetry of septa radial and regularly hexameral. Cycles of septa regular. Septal cycles differ in length, almost not in thickness. First three septal cycles extended to the centre of the calice, later cycles are shorter.
Septa
of younger cycles often and regularly connected to septa of preceeding ones. Septal upper border smooth (probably due to preservation), lateral face with numerous fine thorns, inner margin unknown. Pali or paliform lobes absent. Costae present. Costae ornamented with fine granules on their surface. Synapticulae present, irregularly distributed. Columella composed of a group of isolated granules. Endotheca unknown. Wall present, compact, structure unknown.
REMARKS
For the
type
species, only the complete
type
specimen and thin sections of topotypical specimens were available. The structure of the synapticulae and the endotheca could not be observed because no longitudinal thin sections could be prepared and because the
type
specimen is eroded and barely shows ornamentation. Species other than the
type
species show a regular dentation of the septal upper margin and vertical lines of granulae on the septal lateral face corresponding to the dentations. The existence of fulturae is possible, but could not be proven with the available material; they are reported by
Alloiteau (1957: 339)
. The endotheca is absent in species other than the
type
species, although
Alloiteau (1957:339)
claimed it existed. Alloiteau obviously had more material on hand, but it could not be found at the MNHN. The
type
locality has been inaccessible for a long time and does not allow re-collection.
Actinoseris
cannot be assigned to the
Funginellidae
as formerly proposed.
Funginella
d’Orbigny, 1849
was long considered congeneric with the genera of the
Asteroseriidae
assuming that
F
.
neocomiensis
d’Orbigny, 1850
is the
type
species (for instance
Beauvais 1982
: II, 117). Hence, the
type
species is
Cyclolites haueri
Michelin, 1847
, as designated by
Wells (1936)
. The genus cannot be assigned to the
Siderastraeidae
because of the comparably lower density of ornamentation of the lateral faces of septa and lower number of synapticulae.