<document id="FD978BDF3209F949CF14AFCA9B380936" ID-DOI="10.1649/0010-065X-72.3.433" ID-ISSN="1938-4394" ID-Zenodo-Dep="5383463" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630296642708" checkinUser="diego" docAuthor="Gillett, Conrad P. D. T. &amp; Barr, Iain" docDate="2018" docId="03E32325FF81336387BC3C66FD9BFDC7" docLanguage="en" docName="0010-065X-72.3.433.pdf" docOrigin="The Coleopterists Bulletin 72 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-72.3.433" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Proagoderus lanista" docType="treatment" docVersion="4" lastPageNumber="437" masterDocId="FFDA5B5DFF84336687633D48FFE8FFB5" masterDocTitle="New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia" masterLastPageNumber="438" masterPageNumber="433" pageNumber="437" updateTime="1699219109578" updateUser="plazi">
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<mods:title id="F5D4C469B2D3D998EDCE0CDFB81EDBB4">New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia</mods:title>
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<emphasis id="B93E4E21FF81336387BC3C66FE4EFEF6" bold="true" box="[223,422,302,323]" italics="true" pageId="5" pageNumber="437">Proagoderus lanista</emphasis>
(Castelnau, 1840)
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(
<figureCitation id="13718EB6FF81336386193C01FE2DFEEB" box="[378,453,329,350]" captionStart="Figs" captionStartId="3.[197,236,1616,1635]" captionTargetBox="[220,1145,196,1598]" captionTargetId="figure-1@3.[218,1147,194,1600]" captionTargetPageId="3" captionText="Figs. 1–3. 1) Maps depicting the approximate locations of the main sampling sites (numbered and named) in A) Zambia and B) Swaziland; 2) Habitats at sampling localities – A) Savanna-forest at Mbuluzi Game Reserve, Lubombo, Swaziland, March 2016, B) Montane grassland at Malolotja Nature Reserve, Hhohho, Swaziland, March 2016, C) Miombo woodland, 30 km SSW of Kitwe, Copperbelt Province, Zambia, December 2011; 3) Newly recorded species of scarabaeine dung beetles from Swaziland – A) Copris fidius, B) Catharsius tricornutus, C) Proagoderus lanista. All photographs by Conrad P. D. T. Gillett." figureDoi="http://doi.org/10.5281/zenodo.5383465" httpUri="https://zenodo.org/record/5383465/files/figure.png" pageId="5" pageNumber="437">Fig. 3C</figureCitation>
)
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Previously recorded from 
<collectingCountry id="F35DD2A3FF81336386B83C3CFDD3FE3C" box="[475,571,372,393]" name="Botswana" pageId="5" pageNumber="437">Botswana</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF813363852C3C3CFD67FE3C" box="[591,655,372,393]" name="Kenya" pageId="5" pageNumber="437">Kenya</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF81336387CE3CC6FEC5FE16" box="[173,301,398,419]" name="Mozambique" pageId="5" pageNumber="437">Mozambique</collectingCountry>
, and 
<collectingCountry id="F35DD2A3FF813363860D3CC6FE07FE16" box="[366,495,398,419]" name="South Africa" pageId="5" pageNumber="437">South Africa</collectingCountry>
(
<bibRefCitation id="EFDBEFC2FF81336385603CC6FD78FE16" author="Ferreira, M. C." box="[515,656,398,419]" pageId="5" pageNumber="437" pagination="5 - 1088" refId="ref3778" refString="Ferreira, M. C. 1972. Os escarab'ideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
; 
<bibRefCitation id="EFDBEFC2FF81336387CE3CE1FEA8FE0B" author="Palestrini, C." box="[173,320,425,446]" pageId="5" pageNumber="437" pagination="1 - 358" refId="ref4033" refString="Palestrini, C. 1992. Sistematica e zoogeografia del genere Onthophagus sottogenere Proagoderus Lansberge. Memorie della Societa Entomologica Italiana 124: 1 - 358." type="journal article" year="1992">Palestrini 1992</bibRefCitation>
).
</paragraph>
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<emphasis id="B93E4E21FF81336387A63C8BFE7BFE6D" bold="true" box="[197,403,451,472]" pageId="5" pageNumber="437">New country record</emphasis>
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for 
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: 
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specimens, 
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, 
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<collectingDate id="EFB04D1BFF81336385633C96FD67FE46" box="[512,655,478,499]" pageId="5" pageNumber="437" value="2015-04-14">14 April 2015</collectingDate>
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, and 
<date id="FFF4B4F3FF81336387B43CB1FE86FDBB" box="[215,366,505,526]" pageId="5" pageNumber="437" value="2016-03-31">
<collectingDate id="EFB04D1BFF81336387B43CB1FE86FDBB" box="[215,366,505,526]" pageId="5" pageNumber="437" value="2016-03-31">31 March 2016</collectingDate>
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. Diurnal. The specimens were found during the late afternoon, active under fresh zebra dung in montane grassland
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.
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Genus 
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<emphasis id="B93E4E21FF81336386723F15FE4FFDC6" box="[273,423,605,627]" italics="true" pageId="5" pageNumber="437">Diastellopalpus</emphasis>
van Lansberge 1886
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