The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications Author Sacchetti, Claudia 0000-0002-3225-3139 claudiasacc@icloud.com Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Ávila, Sérgio P. 0000-0002-3225-3139 claudiasacc@icloud.com text Zootaxa 2023 2023-05-24 5295 1 1 150 http://dx.doi.org/10.11646/zootaxa.5295.1.1 journal article 53396 10.11646/zootaxa.5295.1.1 82286fdc-a858-447c-9980-da2e8985d19c 1175-5326 7965273 F3A52660-70B8-439F-A7A0-F45ADC975EA5 Acteon semistriatus (Férussac, 1822) Plate 6 G 1 -G 2 Voluta tornatilis L.— Brocchi 1814: 643 ( partim , pl. 15, fig. 14 only) [ non Acteon tornatilis ( Linnaeus, 1758 ) ]. * Tornatella semi-striata Férussac 1822: 108 . Tornatella semi-striata Fér. — Grateloup 1838: 267 , pl. 6, figs. 18, 21. 1846 Tornatella semi-striata Fér. —Grateloup ( partim , pl. 11, figs. 20, 21 only). Actaeon Burdigalensis d’Orbigny 1852: 35 , no. 519 [ nom. nov. pro Tornatella semi-striata Férussac, 1822 ( partim )]. Actaeon semistriatus Fér ..—Hörnes 1856: 507, pl. 46, figs. 22, 23. Acteon Clathratus Millet 1854: 155 ( nomen nudum ). Acteon clathratus Millet 1865: 581 . Actaeon tornatilis var. semistriata Ferussac —Fontannes 1880: 233, pl. 12, fig. 12. Actaeon semistriatus (Férrusac) — Benoist 1889: 45 , pl. 3, fig. 7. Actaeon Burdigalensis d’Orb. — Benoist 1889: 52 , pl. 4, fig. 1a. Actaeon semistriatus Fér [ sic ]— Sacco 1897: 33 , pl. 3, figs. 21-23. Actaeon semistriatus var. fusulatior Sacco 1897: 33 , pl. 3, figs. 24, 25. Actaeon semistriatus var. maculata (Bors.) — Sacco 1897: 34 , pl. 3, figs. 26, 26 bis. Actaeon semistriatus var. ligustica Sacco 1897: 34 , pl. 3, figs. 27, 28. Actaeon semistriatus var. perstriata Sacco 1897: 34 , pl. 3, fig. 29. Actaeon semistriatus var. totostriata Sacco 1897: 34 , pl. 3, figs. 30, 31. Acteon semistriatus Defrance — Dollfus et al . 1903: 16 , pl. 34, fig. 14. Acteon semistriatus var. totostriata Sacco — Dollfus et al . 1903: 16 , pl. 34, fig. 15. Actaeon semistriatus Fer. —Friedberg 1928: 538, pl. 35, fig. 10. Acteon semistriatus var. burdigalensis d’Orb. —Peyrot 1932: 154, no. 1375, pl. 11, figs. 36, 45-46, 58-60. Acteon semistriatus burdigalensis d’Orbigny, 1852 —Glibert 1952: 384, pl. 13, fig. 7. Actaeon semistriatus Ferrusac — Glibert 1952b: 139 , pl. 10, fig. 10. Acteon semistriatus (Basterot) — Sorgenfrei 1958: 298 , pl. 65, fig. 216. Acteon laevigatus (Basterot) — Sorgenfrei 1958: 300 , pl. 65, fig. 217 [ non de Basterot, 1825 ]. Acteon ( Acteon ) semistriatus (Ferussac 1822) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 219 , pl. 52, fig. 3. Acteon semistriatus var. burdigalensis d’Orbigny —Kecskemétiné-Körmendy 1962: 93, pl. 11, fig. 9. Acteon semistriatus Ferussac, 1822 — Brébion 1964: 643 , pl. 15, figs. 24-27. Actaeon semistriatus (Ferrusac) — Anderson 1964: 330 , pl. 50, fig. 295. Acteon semistriatus Férrusac, 1822 — Strausz 1966: 467 , fig. 209. Actaeon semistriatus (Ferrusac in Grateloup 1840 )— Rasmussen 1968: 214 , pl. 25, figs. 1, 2 (juvenile). Actaeon semistriatus (Férrusac, 1821 [ sic ])— Nordsieck 1972a: 121 , fig. 205. Acteon semistriatus burdigalensis d’Orbigny, 1852 — Steininger 1973: 448 , pl. 9, fig. 7. Actaeon semistriatus (Ferussac) — Fekih 1975: 140 , pl. 15, fig. 15. Actaeon semistriatus ( Ferrusac, 1822 ) [ sic ]— Martinell 1982c: 226 , pl. 1, figs. 14, 15 [in error Actaeon tornatilis ( Linne, 1766 ) on plate text]. Actaeon ( Actaeon ) semistriatus ( Férrusac, 1822 ) —A.W. Janssen 1984: 364 , pl. 13, fig. 13, pl. 18, fig. 1, pl. 78, fig. 2. Actaeon ( Actaeon ) semistriatus f. burdigalensis d’Orbigny, 1852 —A.W. Janssen 1984: 365 , pl. 78, fig. 3. Acteon semistriatus (Férussac, 1822) — Moths 1989: 154 , pl. 21, fig. 112. Acteon semistriatus (Férussac, 1822) — Cavallo & Repetto 1992: 164 , fig. 469. Acteon semistriatus (Férussac) — Ruiz Muñoz et al . 1997: 188 , pl. 41, figs. 5, 6. Acteon semistriatus burdigalensis d’Orbigny, 1852 — Harzhauser 2002: 125 , pl. 12, fig. 10. Acteon ( Acteon ) semistriatus ( Férrusac, 1822 ) — Schnetler 2005: 128 , pl. 9, fig. 17. Actaeon ( Actaeon ) semistriatus ( Férrusac, 1822 ) and Actaeon ( Actaeon ) semistriatus fa. burdigalensis Orbigny — Wienrich 2007: 768 , pl. 132, fig. 9, pl. 168, figs. 4-7. Acteon semistriatus ( Férrusac, 1822 ) — Chirli & Richard 2008: 79 , pl. 16, figs. 1, 2. Acteon semistriatus ( Férrusac, 1822 ) — Chirli 2013: 22 , pl. 7, figs. 7-12. ‘ Acteon ’ semistriatus ( Férrusac, 1822 ) — Ceulemans et al . 2018: 112 , pl. 7, fig. 1 ( cum syn .). ‘ Acteon ’ semistriatus ( Férrusac, 1822 ) — Landau et al . 2020a: 264 , pl. 9, figs. 1-3. Acteon semistriatus (Ferussac, 1822) — Brunetti 2022 , p. 78 , fig. 183. Santa Maria material examined. Maximum height 7.3 mm width 3.9 mm. DBUA-F 191-9 (1). Malbusca, Santa Maria Island, Azores , Touril Complex, Lower Pliocene. Discussion. The specimen from the Azores is very poorly preserved and missing its spire. Nevertheless, the sculpture of spiral cords just below the suture and on the base of the last whorl are typical for the species. As discussed by Landau et al. (2013: 323 ; 2020a: 264 ) ‘ Acteon ’ semistriatus ( Férrusac, 1822 ) is a highly variable species in both profile with slender and squatter forms coexisting and in its sculpture that is hugely subobsolete mid-whorl on the last whorl but can persist along the entire last whorl in some specimen. Distribution. Lower Miocene: Atlantic (Aquitanian and Burdigalian), Aquitaine Basin, France ( Benoist 1889 ; Peyrot 1932); Proto-Mediterranean Sea (Burdigalian), Colli Torinesi, Italy ( Sacco 1897 ); Paratethys (Aquitanian and Burdigalian), Austria ( Steininger 1973 ; Harzhauser 2002 ). Lower-middle Miocene: North Sea Basin (late Burdigalian-Langhian), Belgium ( Glibert 1952b ), Denmark ( Sorgenfrei 1958 ), Germany ( Anderson 1964 ; Moths 1989 ; Wienrich 2007 ), Netherlands ( Nordsieck 1972a ; A.W. Janssen 1984 ). Middle Miocene: Atlantic (Langhian and Serravallian): Aquitaine Basin, France (Grateloup 1846; Benoist 1889 ; Peyrot 1933), (Langhian), Loire Basin, France (Glibert 1952); Paratethys (Langhian-Serravallian), Austria (Hörnes 1856), Bulgaria ( Kojumdgieva & Strachimirov 1960 ), Hungary (Kecskemeìtineì-Körmendy 1962; Strausz 1966 ), Poland (Friedberg 1928); Proto-Mediterranean Sea (Serravallian), Karaman Basin, Turkey ( Landau et al . 2013 ). Upper Miocene: North Sea Basin (Tortonian), Denmark ( Rasmussen 1968 ; Schnetler 2005 ); Atlantic (Tortonian and Messinian): NW France ( Millet 1854 , 1865 ; Brébion 1964 ; Landau et al. 2020a ), Algarve Basin, Portugal ( Dollfus et al . 1903 ), Seville southwestern Spain ( Cárdenas et al. 2019 ); Proto-Mediterranean Sea (Tortonian), Po Basin, Italy ( Sacco 1897 ). Lower Pliocene: Atlantic, NW France ( Brébion 1964 ; Ceulemans et al . 2018 ), Santa Maria Island, Azores (this paper); Guadalquivir Basin, Spain ( Ruiz Muñoz et al . 1997 ; Landau et al . 2011 ; Brunetti 2022 ); western Mediterranean, northeastern Spain , (Martinell 1982), Roussillon Basin, France (Fontannes 1880); central Mediterranean, Italy ( Sacco 1897 ; Chirli 2013 ), Tunisia ( Fekih 1975 ). Upper Pliocene: western Mediterranean, Estepona Basin (NHMW collection), southern France ( Chirli & Richard 2008 ); central Mediterranean, Italy ( Sacco 1897 ; Cavallo & Repetto 1992 ).