Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)
Author
Santos, Daubian
https://orcid.org/0000-0003-1220-1267
Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil
daubians@gmail.com
Author
Santos, Rodrigo dos Reis
Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil
Author
Ribeiro, Guilherme Cunha
Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil
text
Arthropod Systematics & amp; Phylogeny
2022
2022-09-12
80
439
494
http://dx.doi.org/10.3897/asp.80.e83035
journal article
http://dx.doi.org/10.3897/asp.80.e83035
1864-8312-80-439
62FFB94CEBF441639F22881435EFC37C
3F4E21B524465471831D09738C2304A6
3.3.22.
Amphineurus (Rhamphoneurus) rutristylus Alexander, 1968
Amphineurus (Rhamphoneurus) rutristylus
Alexander, 1968: 30 (description, fig. 18: male terminalia).
Material examined.
Holotype
:
♂
, Chile, Valdivia, Enco [
39°53′S
72°8′W
],
1400-1600 m
,
6-III-1955
,
Pena
,
Holotype
12226 (USNM) [part of specimen pinned and part in slide prepared by C.P. Alexander]. -
Additional material
:
CHILE
.
1 ♂
and 1 [sex unknown],
Arauco
,
Nahuelbuta
[
37°46′S
72°59′W
], 13/
20-II-1958
,
Pena
(USNM); 1 [sex unknown],
Caramavida
[
37°40′S
73°20′W
],
1000-1300 m
, 1/
6-I-1954
,
Pena
(USNM); 1 [sex unknown], Lonquimay West, Cordillera las Raices [
38°31′S
71°28′W
],
1500 m
,
25-XII-1968
,
Pena
(USNM)
;
1 ♂
,
Lonquimay West
,
Cordillera
las
Raices
[
38°31′S
71°28′W
],
1500 m
,
25-XII-1968
,
Pena
(USNM)
*.
Diagnosis.
This species is distinguished by a dark spot near R2 and m-cu inserted at fork of bM. Furthermore this species is distinguished by the male terminalia with the lobule with two rounded margins forming heart shape, sheath of aedeagus is straight but without spurs and the mesal lobes are symmetrical, with slender, bare and slightly curved tips.
Redescription.
Wing length 6.87 mm, width 2.00 mm. -
Coloration
: General coloration dark brown. Rostrum and palpus dark brown. First segment of antenna light yellow, remainding segments dark yellow. Head brown. Eyes black. Thorax yellowish-brown, pleura light brown. Scutum without stripes. Halter brownish-yellow. Coxae brownish yellow and legs dark brown. Abdomen dark brown. -
Head
(Fig.
27A
): Rostrum bifurcated with long appendices; first palpal segment slightly shorter than terminal segment; scape thicker than pedicel, terminal flagellomere as long as penultimate, antenna with numerous setae. -
Thorax
(lateral view Fig.
27D
, dorsal view Fig.
27E
): Thorax with several long setae. Anatergite taller than katatergite. Halter with dilated knob. Wing (Fig.
27C
) almost clear of markings, except for marking covering R2; veins near bM (Fig.
27B
) faded; m-cu inserted at fork of bM; M1+2 shorter than basal deflection of M1; R2+3+4 longer than R2+3. -
Male terminalia
(Fig.
27F
): Male tergite IX faintly darkened at posterior margin, with wide V-shaped median notch; posterolateral lobes nearly triangular; ventral branch slender, with concentration of setae on distal tip. Dorsal branch of gonocoxite shorter than half length of ventral branch. Lobe of gonostylus with heart-shaped lobule, more than twice as wide as lobe of gonostylus. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved, oblong-shaped. Mesal lobes of gonocoxite symmetrical, both with narrow, elongate, straight extremity. Sheath of aedeagus straight, slightly bent near tip, darkened at distal one-third.
Figure 27.
Amphineurus (Rhamphoneurus) rutristylus
Alexander.
A
head (dorsal view);
B
detail of wing;
C
wing;
D
thorax (lateral view);
E
thorax (dorsal view);
F
male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations:
aed sh
, sheath of aedeagus;
anatg
, anatergite;
anepm
, anepimeron;
anepst
, anepisternum;
aprn
, anteropronotum;
cgonst
, clasper of gonostylus;
cx
, coxa;
goncx
, gonocoxite;
hlt
, halter;
k
, knob of halter;
kepm
, katepimeron;
kepst
, katepisternum;
ktg
, katatergite;
l ms
, left mesal lobe of gonocoxite;
lgonst
; lobe of gonostylus;
mr
, meron;
mtanepst
, metanepisternum;
mtepm
, metepimeron;
mtg
, mediotergite;
mtkepst
, metakatepisternum;
mtn
, metanotum;
p
, posterior basalare;
patg
, paratergite;
pprn
, postpronotum;
presct
, prescutum; r
ms
, right mesal lobe of gonocoxite;
sct
, scutum;
sctl
, scutellum;
t9
, male tergite IX.
Remarks.
Some specimens of this species were misidentified by C.P. Alexander as
A. (R.) insanus
Alexander. This species resembles
A. (R.) glabristylatus
Alexander but differs mainly in the attachment of m-cu, wing markings, shape of male tergite IX and mesal lobe.