New records of Palaeocorystoidea from the Iberian Peninsula and remarks on the taxonomic significance of the posterior margin of Cenomanocarcinus Van Straelen, 1936 and Hasaracancer Jux, 1971 (Decapoda; Brachyura; Cenomanocarcinidae)
Author
Ossó, Àlex
text
Zootaxa
2017
4303
2
273
283
journal article
32522
10.11646/zootaxa.4303.2.7
2a5c89e9-de41-47fc-8d38-c1a10ae82c41
1175-5326
840956
99DB2319-498E-4A8B-BE10-12B53E2B7C9A
Necrocarcinus woodwardii
Bell, 1863
(
Fig. 4
)
Necrocarcinus Woodwardii
Bell 1863
: 20
, pl. 4, figs. 1‒3.—
de
Tribolet
1875:
8l.
—
Woodward
1877
: 3
.—
Forir
1887
: 193
.—
Carter
1898
: 29
, pl. 2, fig. l.—
Lőrenthey
1929
: 300
, fig. 22a.—
Van Straelen
1936
: 36
.—
Förster
1968
: 175
.—
Wright
&
Collins
1972
: 65
‒66, pl. 12, fig. 1‒2.—
Gründel
1974
: 90
, t. 4, fig. 1‒2.—
Feldmann
et al.
1993
: 36
.—
Fraaye
1994
: 264
, fig. 1.—
Schweitzer
et al.
2003
: t. 3.—
Jagt
et al.
2010
: 178
, t. 1.—
Breton
&
Collins
2011
: 147
, fig. 5
A
–B.—
Van Bakel
et al.
2012
: 204
.—
Luque
2014
: 31
.
Locality and horizon.
Racó
de Cortés
, near Orxeta (
Marina Baixa, Alacant
Province,
Spain
).
Stoliczkaia dispar
Biozone
, late Albian.
Material examined and measurements (in mm).
One specimen embedded in marly limestone. MGB
78175:
length= 23,5; width= 25; fronto-orbital width= 10,5.
Description.
Carapace small sized, appears subcircular, slightly wider than long; the specimen has maximum width at mid length of carapace; dorsal regions covered by strong conical upward directed tubercles; cuticle finely granulated. Fronto-orbital margin about 0,60 of maximum carapace width. Front produced with subtriangular rostrum, distally downturned, with one small tooth on each side. Orbits rounded, raised; supraorbital margin with two notches bounding a supraorbital tooth, and with strong outer and inner orbital teeth. Anterolateral margin convex, triangular teeth present, the last one (epibranchial) laterally directed. Posterolateral margin convex; tooth present at mid length. Posterior margin not preserved. Epigastric region depressed medially. Protogastric lobes slightly swollen, two conical tubercles transversally contiguous. Mesogastric lobe subtriangular elongate, reaching the epigastric region; posterior part wider, with three contiguous tubercles the medial one the most prominent; anterior part slightly inflated, slender, with one tubercle at same level as that of the protogastric ones. Metagastric region indistinct Urogastric region subtrapezoidal, inflated, with a central tubercle. Cardiac region swollen, bearing a prominent tubercle, separated from the urogastric region by a weak depression. Intestinal region depressed, not completely preserved. Hepatic region defined by a small tubercle. Epibranchial region defined by two contiguous tubercles at the same level as the posterior ones of the mesogastric lobe; mesobranchial lobe inflated with prominent medial tubercle; metabranchial lobe depressed, tubercle near the posterolateral corner. Cervical groove well defined, dividing meso-metagastric lobe and urogastric lobe. Branchiocardiac groove deeply marked along urogastric and cardiac regions.
Remarks.
Even although is not possible to count the “twenty large tubercles” described by
Wright & Collins (1972: 66)
because the Orxeta specimen is incomplete, at least eighteen of them can be counted. Despite the lateral and posterior margins of this specimens are not well-preserved, and the carapace is somewhat crushed, its overall outline and regions with strong upward directed conical tubercles, match perfectly with the description and figures of
Wright & Collins (1972)
of British specimens or those of
Breton & Collins (2011)
of French specimens. In addition, the Spanish specimen presents well-preserved triangular rostrum with small rostral spines on both sides, as was pointed out in the French samples by
Breton & Collins (2011: 147)
. The occurrence of
N. woodwardii
in the Albian outcrops of the
Racó
de Cortés
in the southeastern of the
Spain
represents, besides being the first record of the species in the Iberian Peninsula, the most meridional record for the species, only known hitherto from the
Upper Albian
and Cenomanian of the
United Kingdom and France
and
Germany
.