A new monotypic genus and new species of lady beetle (Coleoptera: Coccinellidae Coccinellini) from western South America Author Vandenberg, Natalia J. text Zootaxa 2019 2019-12-20 4712 3 413 422 journal article 24572 10.11646/zootaxa.4712.3.7 4482b4cb-e1f0-48a8-a6e5-b5fda6e93bc9 1175-5326 3586422 80BBA2AF-DDE8-456E-B983-3DA4F66A7181 Tribe Coccinellini Latreille, 1807 Neda -group ( sens. Vandenberg 1992 , Vandenberg & Gordon 1996 , Vandenberg 2002 ; composed of Neda , Mononeda , Neoharmonia , the new genus described below, and various species misclassified in Cycloneda Crotch, 1871 ) ( Figs. 1–4 , 11–19 ) Diagnosis. Medium-sized to large Coccinellini ( 3.5–12.4 mm ) with cardioid, circular, slightly elongate oval, or escutcheonal (sub-triangular, heater-shaped) body form; elytral margins narrowly to broadly explanate; and meso-, metafemora not or only slightly projecting beyond elytral margin in dorsal view ( Figs. 1–4 , 11a ,d–14a,d). Male genitalia with apex of basal lobe (=penis guide sens. Ślipiński) bifid or acuminate ( Figs. 11 e–14e, 16); penis ( Fig. 15 ) with outer arm of basal capsule elongate conical or apically flared, resembling bell of trumpet and forming conduit for ejaculatory duct; inner arm of basal capsule (apodeme) short, weakly arcuate; subapical region of penis with twisted or threaded appearance; apex of penis lacking expanded membranous area bearing chitinous struts or spicuoles. Female genitalia ( Figs. 18–19 ) with sperm duct attached to dorsal end of bursa, enclosed in straight or sinuous tubular sheath (=infundibulum auctorum) for at least part of length; sheath rigid or with reduced flexibility. Spermatheca large, with curved cornu; nodulus typically well developed, but reduced in some; accessory gland large, half length of spermatheca or more; bursa membranous, lacking expanded sclerotized plate or reinforcing struts.