Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae)
Author
Maya-Morales, Julieta
Author
Jiménez, María Luisa
text
Zootaxa
2017
4262
1
1
88
journal article
33105
10.5281/zenodo.571683
5ab1c651-825a-43bc-af00-11b77a792de3
1175-5326
571683
894795FB-ED38-4132-92B9-6AC8C3E41566
Novalena punta
new species
Figure 38
; Map 8
Type
.
Holotype
male
:
MEXICO
:
Veracruz
:
Municipality
of
Las Vigas
,
San Juan
del
Monte
,
2400 m
,
15.IV– 17.V.1999
,
Q. Santiago
and
J. Márquez
(CNAN-T0949), deposited at
CNAN
.
Etymology
. The specific name is a noun in apposition referring to the shape of tegular median process of male pedipalp in ventral view, ‘
punta’
, which means ‘
pointed’
.
Diagnosis
. The males of
N. punta
differ from
N. perote
and
N. valdezi
in having the tegular median process exceeding the length of conductor (
Fig. 38
a). The females differ from
N. comaltepec
in having the atrium withouth sclerotized tips on posterior margin (
Fig. 38
e), from
N. perote
and
N. popoca
in having the atrium strongly procurved, and from
N. valdezi
in having the atrium occupying less than a quarter of the plate length (
Fig. 38
c).
Description
.
Male
(
holotype
): Carapace light brown. Chelicerae dark brown, condyles light brown. Endites and labium dark brown with white tips. Sternum dark brown. Legs yellow with three rings on femur and one on patella. Specimen without opisthosoma. Carapace length 2.05, width 1.29, cephalic region width 0.82, ocular region width 0.42. Eye diameter: AME 0.06, ALE 0.12, PME and PLE 0.1. Distance between eyes: AME-AME 0.04, AME-ALE 0.04, AME-PME 0.08, ALE-PLE 0.04, ALE-ALE 0.17, PME-PME 0.08, PME-PLE 0.06. Clypeus height 0.12. Chelicerae with three retromarginal teeth; basal segment length 0.85, fang length 0.36. Labium wider than long (0.39/0.24). Endites almost straight (distance at their base compared at their tips 0.39/ 0.24). Sternum longer than wide (1.09/0.94). Leg lengths: I—femur 1.48/ patella-tibia 1.86/ metatarsus 1.33/ tarsus 1.05; II—1.43/ 1.71/ 1.14/ 0.95; III—1.43/ 1.52/ 1.19/ 0.9; IV—1.81/ 2.14/ 1.81/ 1.19. Spination: Femur dorsal
I— 1-2
-3/
II—1-2
-3/
III—1-2
-3/
IV—1-2
-3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0;
II—1-1
/ 1/ 0;
III—1-1
/ 1/ 1;
IV—1-1
/ 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0/ 1-1-2/ 1-1-0/ 0;
III—1-1
- 0/ 1-1-2/ 1-1-0/ 1-1-0;
IV—1-1
-0/ 1-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 1-0-1/ 1-0-1; II—0/ 2-2-1/ 0-1-1/ 0- 0-1;
III—2-1
-2/ 2-2-2/ 0-1-1/ 0-0-1;
IV—2-1
-2/ 2-2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus:
I—5
,
II— 5
,
III—5
,
IV—5.
Number of dorsal spines on pedipalp: femur 3, tibia 4. Cymbium length 0.71, width 0.4. Embolus curved; tegular median process with two tips, the mesal one twice the length of ectal one and exceeding the length of conductor; dorsal projection of RTA notched and longer than ventral projection (
Fig. 38
a, b).
Female
(
paratype
) (CASENT 9048866): Carapace light brown, white feathery scales present. Chelicerae and condyles dark brown. Endites and labium light brown with white tips. Sternum dark brown. Legs orange with three rings on femur, one on patella, and two on tibia. Opisthosoma light brown with diffuse black spots. Spinnerets light brown with diffuse black spots. Total length 4.25. Carapace length 2.05, width 1.24, cephalic region width 0.58, ocular region width 0.3. Eye diameter: AME 0.06, ALE 0.1, PME 0.08, PLE 0.12. Distance between eyes: AME- AME 0.04, AME-ALE 0.04, AME-PME 0.13, ALE-PLE 0.02, ALE-ALE 0.19, PME-PME 0.08, PME-PLE 0.04. Clypeus height 0.13. Chelicerae with two retromarginal teeth; basal segment length 0.79, fang length 0.3. Labium wider than long (0.36/0.27). Endites slightly convergent (distance at their base compared at their tips 0.36/0.18). Sternum longer than wide (1.1/0.97). Opisthosoma longer than wide (2.38/1.43). ALS separated by their basal diameter (0.19/0.19), PLS with distal segment slightly shorter than basal segment (0.19/0.21). Leg lengths: Ifemur 1.43/ patella-tibia 1.71/ metatarsus 1.05/ tarsus 0.81; II—1.33/ 1.57/ 0.95/ 0.81; III—1.19/ 1.43/ 1/ 0.67; IV—1.67/ 2/ 1.57/ 0.9. Spination: Femur dorsal
I—1-1
-2/
II—1-1
-2/
III—1-1
-2/
IV—1-1
-2; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0;
II—1-1
/ 1/ 0;
III—1-1
/ 1/ 1;
IV—1-1
/ 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 0- 1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0;
III—1-1
-0/ 1-1-2/ 1-1-0/ 0-1-0;
IV—1-1
-0/ 1-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-0/ 0; II—0/ 2-2-2/ 0-1-1/ 0-0-1;
III—2
-0-2/ 1-1-2/ 0-1-1/ 0-1-1;
IV—2
-0-2/ 1-2-2/ 0-1- 1/ 0-1-1. Number of trichobothria on tarsus:
I—4
,
II—4
,
III—4
,
IV—4.
Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (0.5/0.33); atrium superficial and procurved; anterior hyaline spurs separated by slightly more their basal width (
Fig. 38
c, e). Internal genitalia: copulatory ducts curved in lateral direction, primary spermathecae curved and adjacent (
Fig. 38
d).
FIGURE 38
.
Novalena punta
sp. nov.
: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S1, primary spermathecae; S2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: 0.25 mm.
Variation
. Males (n=2): carapace length 2.05–2.14; patella-tibia I length 1.86–1.95.
Additional material examined
.
Paratypes
:
MEXICO
:
Veracruz
: same locality as holotype,
17.V– 16.VI.1999
,
Q. Santiago
and
J. Márquez
,
1 ♂
(CNAN-T0950); Parque Nacional Cofre de
Perote
,
16.1 km
S
Las Vigas
,
2941 m
,
24.VIII.1967
, R.E. Leech, 1 ♀ (CASENT 904 8866).
Distribution
.
MEXICO
:
Veracruz
(Map 8).