Tokunagaia Saether from China (Diptera: Chironomidae: Orthocladiinae) Author Liu, Yuedan Author Wang, Xinhua text Zootaxa 2006 2006-04-24 1183 43 56 journal article 27045 10.5281/zenodo.2645981 76598eec-0e1e-49ec-8851-641649512277 1175-5326 2645981 C8281FFA-FA6F-420E-8F33-F98AD109F3E6 Tokunagaia subulata new species ( Figs. 13–15 ) Type material Holotype male. CHINA : Jilin Province , Antu County , Changbai Mountain , ( 43°07’N , 128°54’E ), 1700 m a.s.l. , 30.iv.1994 , sweep net , X. Wang (BDN no. 01863) . Paratype : 1 male as holotype . Etymology From Latin, subula , awl­shaped and – atus , provided with, referring to the pointed anal point. Diagnostic characters The male differs from other described species of the genus by having a comparatively large and dark body, a small triangular anal point, long crista dorsalis and a rectangular inferior volsella. This species is similar to T. kinkaensis ( Sasa, 1989 ) , but it is readily separable by the following characters (characters of T. kinkaensis in parentheses): Eyes with wedge­shaped dorsomedian elongation (without); R 2+3 separate (fused with R 4+5 ), with 6 strong acrostichals (without); ground color of thorax dark brown to black (pale yellow). Description Male (n = 2) Total length 2.78–3.23 mm . Wing length 2.28–2.51 mm . Total length/wing length 1.22–1.29. Wing length/length of profemur 2.51–2.30. Coloration [slide mounted material]: body largely dark­brown. Head. AR 1.18–1.22. Temporal setae 6–13; including 0–3 inner verticals, 1–4 outer verticals and 5–6 postorbitals. Clypeus with 8 setae. Tentorium 123–127 µm long. Palpomere lengths (in µm): 27–29, 33–37, 115–117, 88–102, 138–145. Thorax. Antepronotum with 4–6 lateral setae, acrostichals 6, dorsocentrals 7, prealars 3–4. Scutellum with 4–5 setae. Wing ( Fig. 13 ). Anal lobe developed. VR 1.06. R 2+3 ends 1/4 distance between R 1 and R 4+5 . Costal extension 33 µm long. R with 8 setae, R 4+5 with 6 setae, remaining veins bare. Squama with 11 setae. Legs. Spur of front tibia 55–60 µm long, without denticles. Spurs of mid tibia 38–40 µm and 20–25 µm long, of hind tibia 60–68 µm long, all with weak denticles. Hind tibial comb with 14 setae. Two pseudospurs present on ta 1 ­ta 2 of mid and hind legs. Lengths and proportions of legs as in Table 4. Hypopygium ( Fig. 14 ). Anal point small with pointed apex, 13–25 µm long. Tergite IX with 10–12 long setae. Phallapodeme 58–69 µm long; transverse sternapodeme distinctly arcuate, 95–105 µm long, with oral projections. Virga 9–11 µm long, composed of cluster of 7–8 spines. Gonocoxite 168–198 µm long, inferior volsella rectangular, with more than 15 long setae. Gonostylus ( Fig. 15 ), 98–100 µm long, crista dorsalis present, megaseta 8–10 µm long. HR 1.71–1.98; HV 2.78–3.04. Female and immatures: Unknown. Distribution The specimens were collected with a sweep net flying over snow in early spring in a mountain area in the Jilin Province in Northeast China (Palaearctic Region). Tokunagaia unicentrata new species ( Figs. 16–18 ) Type material Holotype male. CHINA : Sichuan Province , Yajiang County , ( 30°02’N , 101°01’E ), 3750 m a.s.l. , 9.vi.1996 , sweep net , X. Wang (BDN no.12258) . Paratypes : 1 male , Sichuan Province , Kangding County , ( 30°04’N , 101°57’E ), 3700 m a.s.l. , 15.vi.1996 , light trap , X. Wang ; 1 male , Sichuan Province , Daocheng County , ( 29°01’N , 100°18’E ), 3750 m a.s.l. , 20.iv.1988 , sweep net , X. Wang . Etymology From Latin, unus , single, centrum , spur and – atus , provided with, referring to the single spur on mid and hind legs. Diagnostic characters The male differs from other known species of the genus, including the similar T. scutellata (Brundin) , by having a single spur on mid and hind tibiae and a strongly sclerotized virga. Description Male (n = 3) Total length 2.88–3.21, 3.05 mm . Wing length 2.25–3.37, 2.81 mm . Total length/wing length 1.28–1.35, 1.31. Wing length/length of profemur 2.55–3.66, 3.11. Coloration [slide mounted material]: head black, thorax reddish brown with dark brown vittae and preepisternum, wings light brown, abdominal segments and legs brown. Head. AR 0.46–0.48, 0.47. Temporal setae 5–6, 6; including 1 inner vertical, 3 outer verticals, and 1–2, 2 postorbitals. Clypeus with 5–6, 6 setae. Tentorium 128–133, 131 µm long. Palpomere lengths (in µm): 31–33, 32; 20–51, 51; 102–112, 106; 88–90, 89; 128–133, 131. Thorax. Antepronotum with 1–2, 2 setae; acrostichals 6–7, 7; dorsocentrals 4–6, 5. Scutellum with 1–2, 2 setae. Wing ( Fig. 16 ). Anal lobe well developed. VR 1.13–1.28, 1.20. R 2+3 ends 1/3 distance between R 1 and R 4+5 . Costal extension 42–52, 47 µm long. Brachiolum 0–1, 1 setae; R with 8–9, 9 setae; R 4+5 with 4 setae; remaining veins bare. Squama with 7–10, 9 setae. Legs. Spur of front tibia 40–45, 39 µm long, without denticles. Spurs of mid tibia 29–33, 31 µm long; of hind tibia 38–53, 46 µm long; both with weak denticles. Hind tibial comb with 10–11, 11 setae. Two pseudospurs present on each of ta 1 and ta 2 of mid and hind legs. Lengths and proportions of legs as in Table 5. Hypopygium ( Fig. 17 ). Tergite IX with 10–12, 11 long setae. Phallapodeme 77–83, 80 µm long; transverse sternapodeme arcuate, 90–120, 105 µm long, with oral projections. Virga dark brown, well sclerotized, 10–14, 12 µm long, composed of cluster of 7–8, 8 spines. Gonocoxite 192–200, 196 µm long; inferior volsella oval, with more than 15 setae. Gonostylus ( Fig. 18 ), 78–82, 80 µm long; megaseta 8–9, 9 µm long. HR 2.43–2.46, 2.45; HV 3.69–3.91, 3.80. Female and immatures: Unknown. Distribution The specimens were collected with sweep net in a subtropical area in the Sichuan Province in Southwest China (Oriental Region).