Revision of Menippus (Coleoptera: Chrysomelidae: Galerucinae) of Taiwan and Menippus dimidiaticornis species group with a new generic synonymy Author Lee, Chi-Feng Author Bezdĕk, Jan Author Suenaga, Haruki text Zootaxa 2012 2012-08-17 3427 1 16 journal article 44535 10.11646/zootaxa.3427.1.1 97ad468b-42f8-40b3-9494-173836944acb 1175-5326 210575 Menippus sericea (Weise) , status revived and new combination ( Figs 5, 6 , 17 , 25 , 32 , 40, 41 , 44 ) Galerucella sericea Weise, 1889: 622 ; Kimoto, 1979: 464 (synonymized with Pyrrhalta dimidiaticornis ). Menippus canellinus : Chûjô, 1935: 161 ( Taiwan ) . Pyrrhalta sericea : Gressitt & Kimoto, 1963: 464 ; Kimoto, 1969: 24 . Issikia dimidiaticornis : Kimoto, 1989a: 248 ; Kimoto, 1991: 9 . Male. Length 8.0– 8.7 mm . Color ( Figs 5–6 ): brown to dark brown; antennae blackish brown except two basal antennomeres; legs brown. Sculpture: head, pronotum and elytra with small, extremely dense punctures. Antenna short, reaching mid-point of body, all antennomeres elongate, relative length of antennomeres II–XI about 1.0: 1.3: 1.5: 1.5: 1.5: 1.5: 1.5: 1.4: 1.3: 1.4. Pronotum widest at apical 2/5, gradually narrowed towards base; pronotal disc with pair of circular depressions, with shallow depression medially, and transverse depression along hind margin laterally; hind margin slightly medially and laterally concave. Prosternal process reduced to a sharp keel between coxae, parallel-sided from base to apex; procoxal cavities almost closed, long thin hypomeral lobes almost reaching apex of posternal process. Elytra elongate, about 1.5X longer than wide; dorsally flat. Apex of last ventrite with a shallow concave excision. Penis wide and weakly curved in lateral view, abruptly constricted towards tip ( Fig. 17 a); in dorsal view apex slightly mucronate, extremely asymmetric; gonopore directed laterally ( Fig. 17 b). Apex of endophallic sclerite broad, margin smooth without tooth ( Fig. 17 b). Female. Length 8.5–10.0 mm. Similar to male. Apex of last ventrite truncate. Sternite VIII ( Fig. 25 ) medially weakly sclerotized, apex truncate and slightly concave at middle, with dense extremely long setae along apical margin, spiculum long. Spermathecal receptaculum strongly swollen ( Fig. 32 ), abruptly narrowed between receptaculum and pump, pump narrow and curved, spermathecal duct slightly projecting into receptaculum. Gonocoxae ( Figs 35, 36 ) separated, triangular, apex with one or two long setae. Diagnosis. Menippus sericea is similar to M. nepalensis sp. nov. and M. gressitti sp. nov. with the extremely asymmetric penis and prominent tectum ( Figs 12, 16, 17 ). This species is characterized by the smooth apex of the endophallic sclerite of the penis ( Fig. 17 b). Type material examined. Lectotype ♀ of Galerucella sericea Weise , here designated, with labels: “Szetchňan (= Sichuan) 1885 G.. Patanin (white label, printed) / sericea ns (white label, handwritten) / ex. Coll. J. Weise (white label, printed) / SYNTYPUS Galeruella sericea Weise, 1889 labeled by MNHUB 2011”. Other material examined ( 126 specimens ). CHINA : 1♀, Fujian, Ziyungdonshan, NW slopes, 700–1100 m , 25°46’N , 117°20’E , 29.IV.2008 , leg. J. Turna ( RBNN ); 1♀, Sichuan, Gingchen-shan (Umg. Heavenly Old Village), 1000–1300m , 18–20.VI. 1995 , leg. Heinz ( MNHUB ); 233, 2♀♀, Sichuan, Mt. Emei, 1000–2000 m , 6.VI.1997 , leg. M. Ouda ( JBBC ); 1♀, Hunan, Guidong env., 26°04’N , 113°56’E , 26–31.V.1994 , leg. Sausa & Jendek ( FKKC ); TAIWAN : 2♀♀, Shinchiku (= Hsinchu city), 1-30.VII.1918 , leg. J. Sonan ( TARI ); 13, 2♀♀, Hsinchu, Mamei, 18.V.2008 , leg. S.-F. Yu ( BPBM ); 13, Hsinchu, Tahunshan, 1.III.2009 , leg. S.-F. Yu ( TARI ); 13, 1♀, Hsinchu, Wuchihshan, 14.V.2008 , leg. S.-F. Yu ( HSHJ ); 433, 1♀, same locality, 15.V.2008 , leg. H. Lee ( JBBC ); 1♀, Hualien, Huojanting, 12.IX.2007 , leg. Y.-F. Hsu ( TARI ); 1♀, Ilan, Fushan, 3.IV.2008 , leg. M.-H. Tsou ( TARI ); 233, 2♀♀, Kaohsiung, Tengchih, 29.VIII.1996 , leg. W. I. Chou ( NMNS ); 1♀, same locality, 2–5.VI.2008 , leg. C.-F. Lee ( TARI ); 1♀, Nantou, Takeya (= Chienchi), 8.VII.1940 , leg. M. Chûjô ( TARI ); 333, Nantou, Chunyung, 24–25.III.1993 , leg. W. T. Yang ( NMNS ); 1♀, same locality, 8–10.III. 1999 , leg. C. S. Lin & W. T. Yang ( NMNS ); 1♀, same locality, 12.III. –9.IV.2002 , leg. C. S. Lin & W. T. Yang ( NMNS ); 1♀, same locality, 10.VIII. –8.IX.2004 , leg. C. S. Lin & W. T. Yang ( NMNS ); 2♀♀, Nantou, Fenghuangku, 15.IV.1994 , leg. M. L. Chan & W. T. Yang ( NMNS ); 13, same locality, 8–9.III.1995 , leg. W. T. Yang ( NMNS ); 13, Nantou, Hoawanta, 30.IV.1992 , leg. W. T. Yang ( NMNS ); 1♀, Nantou, Nanshanchi, 17.VI.1965 , leg. B. S. Chang ( NMNS ); 13, same locality, 4.IV.2010 , leg. Y.-T. Wang ( TARI ); 1♀, Nantou, Huishun, 11.VII.1992 , leg. Shiau & Yang ( NMNS ); 13, Nantou, Lienhuachih, 22.III.2009 , leg. Uika Ong ( TARI ); 4♀♀, Nantou, Shihtyutou, 2.VIII.1994 , leg. C. C. Lo ( NMNS ); 1♀, Nantou, Tungpu, 1200m , 18–23.XI.1981 , leg. T. Lin & W. S. Tang ( TARI ); 1♀, same locality, 19–23.VII.1982 , leg. L. Y. Chou & T. Lin ( TARI ); 333, 3♀♀, same locality, 20–24.VI.1983 , leg. K. C. Chou & C. Y. Wong ( TARI ); 13, 1♀, same locality, 23–27.VII.1984 , leg. K. C Chou, & C. H. Yang ( TARI ); 1♀, same locality, XI.1985 , leg. K. S. Lin ( MCZC ); 13, Nantou, Wanfengtsun, 10.VII.2007 , leg. W.-T. Liu ( TARI ); 13, same but with “ 4.X.2007 ” ( TARI ); 233, 2♀♀, same but with “ 2.IV.2008 ” (1♀: MNHUB ; 1♀: TARI ); 1♀, same but with “ 7.VII.2008 ” ( TARI ); 13, same but with “ 9.VII.2008 ” ( TARI ); 1♀, Nantou, Musha (= Wushe), 1150m , 18.IV. –15.VI.1919 , leg. T. Okuni ( TARI ); 13, same locality, 26–28.VIII.1981 , leg. L. Y. Chou & S. C. Lin ( TARI ); 13, 1♀, same locality, 19–22.IV.1983 , leg. K. C. Chou & S. P. Huang ( TARI ); 13, same locality 28–29.IV.1993 , leg. C. S. Lin & W. T. Yang ( NMNS ); 1♀, Pingtung, Shuangliu, 4.V.2005 , leg. J.-F. Tsai ( TARI ); 13, Pingtung, Tahanshan, 20.VII.2007 , leg. S.-F. Yu ( TARI ); 1♀, same locality, 8.V.2009 , leg. Uika Ong ( TARI ); 13, 1♀, same locality, 18.V.2009 , leg. M.-L. Jeng ( TARI ); 2♀♀, same locality, 28.VI.2009 , leg. I.-T. Chung ( TARI ); 13, same locality, 1.VIII.2009 , leg. Uika Ong ( TARI ); 13, same but with “ 6.III.2010 ” ( TARI ); 3♀♀, same but with “ 10.IV.2010 ” ( TARI ); 1♀, same locality, 14.IV.2011 , leg. J.-C. Chen ( TARI ); 333, same locality, 14.VIII.2011 , leg. Y.-T. Wang ( TARI ); 1♀, Pingtung, Wutai, 1.IV.2010 , leg. Uika Ong ( TARI ); 13, 1♀, Taichung, Chiapaotai, 750m , 14–18.X.1980 , leg. K. S. Lin & C. H. Wang ( TARI ); 233, Taichung, Wushihkeng, 19.III.2008 , leg. H.-T. Shih ( TARI ); 13, Tainan, Meiling, 12.III.2011 , leg. M.-L. Jeng ( TARI ); 1♀, Taipei, Chutzuhu, 15.VI.2008 , leg. S.-F. Yu ( TARI ); 13, 1♀, Taipei, Peitou, 5.IV.2008 , leg. M.-H. Tsou ( TARI ); 333, Taipei, Manyuehyuan, 9.IX.2011 , leg. H. Lee ( TARI ); 1♀, Taipei, Wulai, 15.VII.1969 , leg. Y. Hori ( EUMJ ); 13, same locality, 7.IV.2002 , leg. Gy. Fábián & O. Merkl ( JBBC ); 13, Taipei, Yangmingshan, 5.VIII.1969 , leg. Y. Hori ( EUMJ ); 433, 2♀♀, same locality, 5.IV.2008 , leg. M.-H. Tsou ( TARI ); 13, same locality, 17.VIII.2008 , leg. S.-F. Yu ( TARI ); 1♀, Taitung, Guanshan, 31.X.2009 , leg. P.-F. Wang ( TARI ); 13, Taitung, Lichia, 2.VI.2009 , leg. Uika Ong ( TARI ); 13, Taitung, Litao, 26.IX.2007 , leg. J.-F. Tsai ( TARI ); 1♀, Taitung, Tulanshan, 8.XII.20092.II.2010 , leg. W. T. Yang & K. W. Huang ( NMNS ); 233, same but with “ 14.IX. –23.XI.2010 ” ( NMNS ); 333, 3♀♀, TAIWAN : Taoyuan, Mingchi, 27.IV.2008 , leg. M.-H. Tsao (= Tsou) ( TARI ). 1♀, same but with “ 5.IV.2009 ”; 1♀, Taoyuan, Paling, 800m , 3–5.V.1983 , leg. K. C. Chou & C. C. Pan ( TARI ); 13, Taoyuan, Gaogan (= Sankuang), 16.VII.1939 ( TARI ); 233, 2♀♀, Taoyuan, Tungyanshan, 24.IV.2008 , leg. H. Lee ( RBNN ); 13, 1♀, same but with “ 9.V.2008 ” ( TARI ); 13, same locality, 22.VII.2008 , leg. J.-C. Chen ( TARI ); 13, 1♀, same locality, 17.IV.2009 , leg. H. Lee ( TARI ); 13, 1♀, same locality, 10.V.2009 , leg. M.-H. Tsou ( TARI ). FIGURES 33–41. Gonocoxae of Menippus species. 33. M. asahinai ; 34. M. beeneni ; 35. M. dimidiaticornis ; 36. M. gressitti ; 37. M. gressitti ; 38. M. hsuehleeae ; 39. M. murzini ; 40. M. sericea from Taiwan; 41. M. sericea from China (Hunan). Host plants. Ulmaceae : Celtis sinensis Pers ; Moraceae : Morus australis Poir. , Broussonetia papyrifera (L.) L'Herit. ex Vent.; Rosaceae : Rubus swinhoei Hance ; Betulaceae : Alnus formosana (Burkill ex Forbes & Hemsl.) Makino ; Aquifoliaceae : Ilex formosana Maxim. ; Lythraceae : Lagerstroemia subcostata Koehne ; Fagaceae : Quercus variabilis Bl. Distribution. China (Fujian, Sichuan, and Hunan) and Taiwan . This species is widespread from lowlands to elevations up to 1500 m in Taiwan ( Fig. 44 ). Notes. Taiwanese population was originally recorded as Menippus canellinus by Chûjô (1935) . Kimoto (1969) identified it as Pyrrhalta sericea . Latter Kimoto (1989a , 1991 ) recorded it as Issikia dimidiaticornis due to synonymy. Acutally M. sericea is a distinct species which is different from M. dimidiaticornis . Biological notes. This species is apparently univoltine. Larvae were found in spring. A few adults were collected in light traps