Systematics and biogeography of the New World species of Trichadenotecnum Enderlein (Insecta: Psocodea: ‘ Psocoptera’: Psocidae) Author Yoshizawa, Kazunori Author Aldrete, Alfonso Neri García Author Mockford, Edward L. text Zoological Journal of the Linnean Society 2008 2008-08-31 153 4 651 723 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00398.x journal article 10.1111/j.1096-3642.2008.00398.x 0024-4082 5446195 TRICHADENOTECNUM QUAESITELLUM SP. NOV. Material examined: Holotype . Male. Mexico : Tamaulipas , 5.1 mi W Gómez Farías , 16.vi.1962 , ELM et al. ( ISU ). Figure 21. Male terminalia structures of Trichadenotecnum quaesitum , showing terminalia in lateral view (A), epiproct in posterior view (B), hypandrium in ventral view (C) and phallosome in ventral view (D). Etymology: The specific epithet is taken from quaesitum (= specific epithet of Trichadenotecnum quaesitum ) + - ellus (= Latin diminutive). This species is very similar to T. quaesitum but smaller. Description: Male . Head . White in ground colour; vertical and orbital markings blackish brown; coronal suture black; frontal suture bordered with narrow brown band dorsally; frons with pair of pale brown bands at middle and with pair of circular markings laterally; eye black, IO/D = 0.6; ocelli white, ocellar field black; gena white; antennal socket bordered with brown band; postclypeus with c . eight rows of pale brown spots at middle, fused with each other ventrally forming a large marking, ventro-lateral corner white, lateral margin of postclypeus dark brown, Key to species of the quaesitum group 1. Forewing extensively covered with small spots ( Fig. 17F–I ).......................................................................3 – Forewing not extensively covered with small spots ( Fig. 17C, D )................................................................2 2. Hypandrial median tongue relatively short, about 3¥ longer than apical width; hypandrial right process exceeding left one ( Fig. 21C )............................................................................................. T. quaesitum (Chapman) – Hypandrial median tongue relatively long, about 3.5¥ longer than apical width; hypandrial right arm not exceeding left one ( Fig. 22C ); female unknown................................................................... T. quaesitellum sp. nov. 3. Hypandrial left process long, exceeding apex of right one ( Fig. 23C ). .......................................................... 4 – Hypandrial left process short, not reaching right one ( Fig. 31C ); female unknown ...... T. brevicornum sp. nov. 4. Forewing sparsely spotted, areola postica not covered with large brown marking ( Fig. 17E ) ........................... 5 – Forewing densely spotted; distal half of areola postica covered with large brown marking ( Fig. 17F )......................................................................................................... T. maculatum sp. nov. 5. Clunial lobe membranous ( Fig. 23A ); subgenital plate without narrow membranous notch antero-medially ( Fig. 24A ).........................................................................................................................................6 – Clunial lobe strongly sclerotized ( Fig. 29A ); body of subgenital plate with narrow membranous notch anteromedially extending near base of egg guide ( Fig. 30A ).................................................... T. ericium sp. nov. 6. Forewing proximal band complete in cell CuA ( Fig. 17E ); hypandrial median tongue broad, about 2¥ longer than apical width ( Fig. 23C ); dorsal margin of epiproct lobe with conical median projection ( Fig. 23B ) ...................... ....................................................................................................................... T. cerrosillae sp. nov. – Forewing proximal band narrowly interrupted at median part of cell CuA ( Fig. 17G ); hypandrial median tongue slender, about 2.5¥ longer than apical width ( Fig. 27C ); dorsal margin of epiproct lobe without distinct median projection ( Fig. 27B ).......................................................................................... T. neoleonense sp. nov. dorsal margin without marking; anteclypeus pale brown. Antenna pale brown, pedicel and scape darker. Mouth parts pale brown; 4th segment of maxillary palpus brown. Thorax . Prothorax brown. Mesonotum pale; anterior surface of scutum brown; lateral lobe of scutum with large pale brown marking; scutellum pale brown, median part darker, anterior margin blackish brown; postnotum brown. Metanotum pale brown, with three whitish oval spots on posterior regions of scutal lobes. Meso- and metapleurites brown, with longitudinal white band medially. Legs. Pale brown; middle and hind coxae, basal half of middle femur and entire hind femur brown, distal tip of hind tibia darker. Forewing ( Fig. 17D ). Not extensively covered with tiny spots. Spots in cell a 1 apparent, nearly of equal size. Opposing spots in cell r distinct. Proximal band broad, narrowly interrupted just beyond fork of M + Cu. Distal band distinct. Spot on roof of cell m 3 narrow but distinct. Submarginal spots apparent; spot in cell r 5 elongate. Marginal clouds distinct. Hindwing. Hyaline, cell CuP with faint brownish spots in distal half; veins brown. Abdomen . White, with sparse minute blackish brown spots. Terminalia . Clunial arm ( Fig. 22A ) developed but very weakly sclerotized, not forming free process apically, basally with membranous lobe ventrally covered with short spines. Anterior margin of 8th sternum ( Fig. 22C ) smoothly arched, fusion with hypandrium narrow. Epiproct lobe ( Fig. 22B ) broadened dorsally, wider than long, dorsal margin sinuate, posterior surface slightly concave. Paraproct ( Fig. 22A ); ventro-lateral band short, triangular; distal process long. Hypandrium ( Fig. 22C ): outer margin of left process slightly arched, directed posteriorly; right process projected outward, then strongly bent postero-internally near base, its apex not exceeding outer margin of left process; median tongue long, slightly constricted medially, apical margin slightly hollowed, dorsal keel rounded. Phallosome ( Fig. 22D ): pseudoparameres widely separated, long, longer than basal width, directed dorsally, apical margin rounded with tiny denticles internally. Phallobase widest at anterior third, then narrowing to pointed anterior end. Length . B 2.3; FW 2.8; HW 2.0. Female . Unknown. Distribution: Mexico .