A revision of the genus Saccoglossus (Hemichordata: Enteropneusta: Harrimaniidae) with taxonomic descriptions of five new species from the Eastern Pacific Author C. B. Cameron Author C. Deland Author T. H. Bullock text Zootaxa 2010 2010-05-25 2483 1 22 journal article 32068 10.5281/zenodo.893894 2bb49143-5cbd-42f7-9689-de741f351ff7 1175-5326 893894 Genus Saccoglossus Schimkewitsch, 1892 Dolichoglossus : Spengel, 1893 Type species. Saccoglossus kowalevskii ( Agassiz, 1873 ) Diagnosis. Proboscis usually long; middorsal longitudinal groove may be present. Collar usually very short compared to proboscis. Dorsal interbranchial genital ridges and dorsal gonads absent but lateral extrabranchial genital ridges may be present. Intestinal pores often present. Perihaemal cavities always present. Peribuccal cavities usually present, but not always. Genus most importantly characterized by arrangement of longitudinal muscle fibers of proboscis in several concentric rings ( Fig. 2 A ). Many species favor quiet muddy-sandy flats not too far from the mouth of a bay, living in ‘permanent’ tubes and throwing up low conical mounds of quasi-spiral castings from the anus. Remarks. The genus as revised above would now include the following species: S. apatensis Thomas, 1956 ; S. aulakoeis Thomas, 1968 ; S. bromophenolosus King, Giray & Kornfield, 1994 ; S. horsti Brambell & Goodhart, 1941 ; S. hwangtauensis (Si & Kwang-Chung, 1935) , S. inhacensis (Kapelin, 1936) , S. kowalevskii ( Agassiz, 1873 ) , S. madrasensis Rao, 1957 , S. mereschkowskii ( Wagner, 1885 ) , S. otagoensis ( Benham, 1895 ) , S. pusillus ( Ritter, 1902 , 1929 ), S. ruber Tattersall, 1905 , and S. sulcatus ( Spengel, 1893 ) . S. cambrensis (Brambell, Rogers and Goodhart, 1939 ) , S. serpentinus (Assheton, 1908) and S. ruber ( Tattersall, 1905 ) are synonymous species from the United Kingdom ( Burdon-Jones, 1960 ) and so S. ruber takes precedence. S. ruber characteristics found in this article, including Table 1 , are taken from the more comprehensive taxonomic treatment of Brambell, Rogers and Goodhart (1939) . Five new species collected from the Eastern Pacific are described below.