Parasitic copepods of the family Lernanthropidae Kabata, 1979 (Copepoda: Siphonostomatoida) from Australian fishes, with descriptions of seven new species
Author
Boxshall, Geoff A.
Author
Bernot, James P.
Author
Barton, Diane P.
Author
Diggles, Ben K.
Author
Q-Y, Russell
Author
Atkinson-Coyle, Toby
Author
Hutson, Kate S.
text
Zootaxa
2020
2020-02-17
4736
1
1
103
journal article
24025
10.11646/zootaxa.4736.1.1
c76ec731-0dc9-4fc4-8ea4-d90d90da9438
1175-5326
3669745
970D7D36-6D8C-4463-B9EA-D3B8E191BE72
Genus
Sagum
Wilson, 1913
Remarks:
The genus
Sagum
was established by
Wilson (1913)
to accommodate a new species,
S. flagellatum
Wilson, 1913
, found on the gills of a serranid
Epinephelus adscensionis
(Osbeck, 1765)
caught off
Jamaica
. The new genus was distinguished from
Lernanthropus
by “the rostrum and wings of the cephalothorax, by the long endopods of the third legs, and by the lash-like tips of the fourth legs.” The two female
syntypes
of
S. flagellatum
lacked egg sacs, so the form of the egg sacs was not included in the original generic diagnosis. In his revision of the North American
Dichelesthiidae,
Wilson (1922)
transferred
Lernanthropus angulatus
Krøyer, 1863
into
Sagum
as
S. angulatum
and gave a revised generic diagnosis which included as a key feature, the possession of coiled egg sacs entirely contained within the space enclosed between the dorsal trunk plate and the third and fourth legs.
In their phylogenetic analysis of the lernanthropid genera,
Ho & Do (1985)
grouped
Aethon
,
Norion
and
Sagum
together on the basis of the possession of coiled egg sacs. They distinguished
Aethon
by its highly modified second legs and grouped
Sagum
and
Norion
by the shared possession of bilobate fourth legs with lobes that are lamellate proximally and flagellate distally. Although used by
Wilson (1922)
is his original diagnosis of the genus, this latter character is not robust since at least half of
Sagum
species (see
Table 4
) have tapering bilobate fourth legs rather than legs with flagellate tips on the exopodal and endopodal lobes. The loss of leg 2 was used to characterize
Norion
species whereas the form of leg 3, described as “divided into 3 branches; lamelliform or cylindrical”, was used to diagnose the
Sagum
branch. This latter character is also not robust, applying only to a minority of species. Given the difficulty in using some of these traditional characters, it is clear that the relationships between the currently recognized lernanthropid genera need to be reassessed using a wider range of characters from both sexes.