Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae) Author Espeland 1, Marianne Author Nakahara, Shinichi Author Zacca, Thamara Author Barbosa, Eduardo P. Author Huertas, Blanca Author Marín, Mario A. Author Lamas, Gerardo Author Benmesbah, Mohamed Author Brévignon, Christian Author Casagrande, Mirna M. Author Fåhraeus, Christer Author Grishin, Nick Author Kawahara 1, Akito Y. Author Mielke, Olaf H. H. Author Miller,] acqueline Y. Author Nakamura, Ichiro Author Navas, Vanessa Author Patrusky, Brooke Author Pyrcz, Tomasz W. Author Richards, Lindsay Author Tan, Denise Author Tyler, Stephanie Author Viloria, Angel Author Warren 1, Andrew D. Author Xiao, Lei Author Freitas, André V. L. Author Willmott, Keith R. text Zoological Research 2023 2023-02-15 2023 1 73 journal article 10.1111/syen.12590 bfb878f3-8a74-46d3-a104-36485c32aaba 7909395 Euptychoides Forster, 1964 albofasciata (Hewitson, 1869) inani (Staudinger, [1886]) , comb.n. , was Macrocissia [Andrade et al. (2019) placed this species in Macrocissia , but it does not cluster with the type species of that genus, instead being strongly supported by molecular data as a member of a clade that includes the type species of Euptychoides ] = juani (Staudinger, 1887) , missp., comb.n. , was Macrocissia laccine (C. Felder & R. Felder, 1867) = saturnus (Butler, 1867) , syn.n. [number of pupils in VHW ocelli is variable even between wings of the same individual] pseudosaturnus Forster, 1964