A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae)
Author
Breitkreuz, Laura C. V.
Author
Ohl, Michael
Author
Engel, Michael S.
text
Zootaxa
2016
4063
1
1
66
journal article
37206
10.11646/zootaxa.4063.1.1
9608b695-6310-4c63-bd94-46bda058e6d8
1175-5326
270412
80401ED8-C6BA-4420-9109-854C5CC1E88D
Arpactophilus nere
sp. nov.
(
Figs. 77–78
)
Diagnosis.
Arpactophilus nere
is the only New Caledonian species in the genus in which the clypeus and labrum have two, prominent, pointed long teeth (
Fig. 77
). Other distinctive features are the yellow markings on clypeus and frons (
Fig. 77
) as well as the yellow mesoscutellum.
Description.
FEMALE: Total length
5.4 mm
; forewing length
3.6 mm
.
Body black, with areas of yellow. Yellow: lateral clypeus; labrum (slightly darker); mandible; palpi; scape; pedicel; area on frons close to inner compound eye margin, broad on level of scape, narrow towards vertex (
Fig.
77); area around mandibular base; anterior pronotal margin; dorsal transversal band on pronotum; band on pronotum parallel to ventral pronotal margin; pronotal lobe; area on mesopleuron below pronotal lobe; area on mesopleuron next to lower 2/3 of mesopleural fissure; tegula; anterior part of mesoscutellum (similar to coloration in
Fig. 19
); legs. Flagellum brown. Wings hyaline; pterostigma dark brown.
Head about 0.9 × as long as wide in frontal view. Apical margin of clypeus projecting medially with steep Vshaped incision, apicolateral angles of incision forming two long, pointed teeth (
Fig. 77
). Apical margin of labrum projecting medially and with deep V-shaped incision, apicolateral angles of incision forming two long, pointed teeth that project beyond apex of clypeal margin (
Fig. 77
). Ventral mandibular tooth nearly 1/3 of total mandibular length, almost reaching opposite mandibular base. Palpal formula 4:3. Frons imbricate with sparse setae. Frontal carina present from median ocellus to upper half of clypeus, there slightly elevated. Scape 7.2 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by more than their diameter. Occipital carina interrupted dorsally (
Fig. 6
). Gena imbricate with sparse setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line faintly carinate, posteriorly with bordering short transverse carinulae on hypostomal integument, not angulate.
Mesosoma about 1.8 × as long as wide in dorsal view. Propodeum about 1.0 × as long as wide in dorsal view. Mesosoma imbricate (
Fig. 78
), except dorsal surface of propodeum; lateral surface of propodeum also transversely carinulate; dorsal surface of propodeum coarsely reticulate (similar to pattern in
Fig. 18
). Impressed, but not pitted sulcus present posterior to mesoscuto-mesoscutellar sulcus. Hypersternal sulcus present as concave elongate area close to omaulus. Metafemur 3.5 × as long as wide. Metatibia without differently colored area apically. Pretarsal claws with teeth. Forewing with two submarginal cells; anterior border of submarginal cell II shorter than posterior border.
Hind
wing with five distal hamuli.
Metasoma matt, punctation sparse. Metasomal sternum II without bulge (as in
Fig. 1
). Pygidium with broad row of silk setae apically.
MALE: Difference from female aside from genitalic structures: generally lighter brown; teeth on apical margin of clypeus much smaller and not pointed; mandible tooth shorter. Metasomal sternum VIII slightly narrow.
Remarks.
Arpactophilus nere
is easily identifiable by its prominent pointed teeth of the apical margin of clypeus and labrum (
Fig. 77
). The most similar species is
A. merle
, which mainly differs in the aforementioned structure of the clypeal and labral margins and coloration of the upper frons. See also remarks-section of
A. merle
.
Material examined.
HOLOTYPE
♀: “
NEW
CALEDONIA
11522:
22°02’S
x
166°28’E
,
950m
, Dzumac Road, junction.
26 Jan 2004
. G. Monteith. pyrethrum, trees & logs” [QM].
PARATYPE
1♂
: “
NEW
CALEDONIA
11522:
22°02’S
x
166°28’E
,
950m
, Dzumac Road, junction.
26 Jan 2004
. G. Monteith. pyrethrum, trees & logs” [QM].
Etymology.
The specific epithet is taken from the name
Nerë
, one of the New Caledonian native languages. It is treated as a noun in apposition.