A New Subfamily of Spiders with Grate-shaped Tapeta from Australia and Papua New Guinea (Araneae: Stiphidiidae: Borralinae) Author Gray, Michael R. Author Smith, Helen M. text Records of the Australian Museum 2008 2008-06-25 60 1 13 44 http://dx.doi.org/10.3853/j.0067-1975.60.2008.1493 journal article 10.3853/j.0067-1975.60.2008.1493 2201-4349 5239981 Elleguna major n.sp. Figs 8d , 16a–e Type material. Queensland : HOLOTYPE : ? , KS34765, lower slopes of Mt William , Dalrymple Heights near Eungella , 21°01'S 148°36'E , Apr.1975 , M. Gray & C. Horseman , in log, Rainforest Survey Site 15, notophyll evergreen vine forest . PARATYPES : ! , KS57656, Eungella area , Snake Rd , 4.7 km NW Dalrymple Rd , 21°02'47"S 148°32'17"E , 20 Apr. 1998 , G. Milledge ; ? , KS34585, Dalrymple Heights , 21°04'S 148°35'E , 9 Apr. 1975 , M.R. Gray & C. Horseman , in roadside bank near Rainforest Survey Camp ; ! , KS57653, Eungella NP , Dalrymple Rd , 1.3 km NE Snake Rd junction, 21°04'S 148°35'E , 18 Apr. 1998 , G. Milledge ; ! , KS57654, as KS57653 but 1.7 km ; 2 ?? , 4 !! , KS83622–7, data as for holotype . Diagnosis . Differs from E. minor as follows: male palp with a large ventral conductor process, a basally angled and apically elongated embolus, and a large RDTA; epigynal fossa indistinctly divided by a weak median ridge and lateral fossa margins extended inwards as thin “flanges”. Description Male ( holotype ). BL 6.65, CL 3.22 (range 3.10–3.39), CW 2.33, CapW 1.22, EGW 0.88, LL 0.43, LW 0.43, SL 1.59, SW 1.35. Legs: 1423 (I: 15.58; II: 13.17; III: 11.58; IV: 14.25); ratio tibia I length:CW = 1:0.66. Male palp ( Fig. 16a,b ). Bulb with a reduced, indistinct TL, with a tightly looped sperm duct visible retrolaterally. Embolus a long, curved spine, thick and strongly angled basally just before entering the conductor groove, then tapering and slender distally. Prolateral limb of conductor wide with a triangular flange; retrolateral limb extended as a long, translucent, curved spine supporting the elongate embolus and making a loose loop around the ventral conductor process.VCP very large with a thickened base. TW basal. MA surrounded by a collar-like membraneous area. RDTA a robust, ventrally concave, tapering process. Fig. 16. Elleguna major . ( a,b ), male palp (holotype): a , ventral; b , retrolateral. ( c–e ), epigynum: c , ventral, fl = flange (KS57656); d , ventral, flanges broken (KS83624); e , dorsal—internal genitalia (KS83624). Scale lines 0.5 mm: upper, a,b ; lower, c–e . Female (KS57656). BL 8.37, CL 4.00 (range 3.22–4.00), CW 2.53, CapW 1.88, EGW 1.16, LL 0.61, LW 0.59, SL 1.80, SW 1.51. Legs: 1423 (I: 14.33; II: 12.25; III: 10.58; IV: 13.00); ratio tibia I length:CW = 1:0.69. Epigynum ( Fig. 16c–e ). Fossa longer than wide, weakly divided by a median ridge; anterolateral fossa margins extended as thin, flangelike laminae (“fl” in Fig. 16c ) over the anterolateral fossa floor and copulatory openings (these flanges may be broken off— Fig. 16d ); posterior fossa floor with a pair of adjacent “socket-like” recesses. Copulatory duct openings anterolateral, widely separated. Internal genitalia. Copulatory ducts very short, of moderate width; spermathecae small. Distribution . Known only from Eungella NP area, mideastern Queensland . Etymology . The species name refers to the large size of the male palpal ventral conductor process.