Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan Author Tasaku, Yuto Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. & Corresponding author Author Ono, Hiroki Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan. Author Maruyama, Munetoshi The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan. text Zootaxa 2023 2023-12-12 5383 3 251 296 https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454 journal article 10.11646/zootaxa.5383.3.1 1175-5326 10361564 20988E42-C14D-4878-A716-8CEAB5E5EF92 Paramblopusa borealis (Casey, 1906) [Japanese name: Nise-nokoba-umi-hanekakushi] ( Figs. 21A–D ; 22 ) Amblopusa borealis Casey, 1906: 355 (original description; type locality: Queen Charlotte Island ( Massett )). Paramblopusa borealis Ahn & Ashe, 1996: 151 (redescription); Ahn & Maruyama, 2000: 364 (characters). Material examined. Japan : Hokkaidô : 1 male , 1 female , 8 unsexed, Otaru-shi , Oshoro , 18. IV . 2020 , S. Hori ; 1 unsexed, Otaru-shi , Shukutsu , 4. IV . 2020 , Y. Tasaku (cYT) ; 1 male , Hakodate-shi , Toyosaki-chô , 12. IV . 2020 , K. Mitsuhashi (cYT) ; 2 males , 2 females , 8 unsexed, 12. IV . 2020 , D. Sumikawa & Y. Tasaku (cYT); 1 male , 9 unsexed, Hamanaka-chô , Mochirippu , Mochirippu-numa , 17. IX. 2020 , Y. Tasaku (cYT); 180 unsexed, ditto, 12. IX. 1999 , M. Maruyama ( KUM ); 1 unsexed, Betsukai-chô , Odaitô , 23. VII . 2020 , Y. Tasaku (cYT) ; 1 male , Akkeshi-chô , Akkeshi-ko , 3. VIII. 2022 , Y. Tasaku (cYT); 1 unsexed, Nemuro-shi , Onnetô , 18. IX. 2020 , Y. Tasaku (cYT); 10 unsexed, Nemuro-shi , Shunkunitai , 18. IX. 2020 , Y. Tasaku (cYT); 24 unsexed, ditto, 11. IX. 1999 , M. Maruyama ( KUM ); 5 unsexed, Samani-chô , Horoman , 2. V. 2023 , D. Sumikawa (cYT) . Diagnosis. This species can be distinguished from other liparocephaline species by the combination of the following characteristics: body slender, head almost circular, abdominal segments III–VII almost parallel-sided ( Fig. 21A ); eyes present; labial palpus 3-segmented and segment II thinner than I ( Fig. 22C ). Habitat. Adults were collected on gravel beaches and under stones on mudflats. Distribution. Japan : Hokkaidô ; Canada ; USA : Alaska , Oregon , Washington .