Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species Author Criscione, Francesco Author Law, Margot Louisa Author Köhler, Frank text Zoological Journal of the Linnean Society 2012 Zool. J. Linn. Soc. 2012-11-26 166 4 689 722 http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x journal article 10.1111/j.1096-3642.2012.00863.x 0024-4082 5409181 AC086999-76A8-4F93-9A51-5B5B213A50E6 EXILIGADA FLORAEVALLIS SP. NOV. ( FIGS 4E , 5D , 9D–F , 11 ) Holotype : AUSTRALIA , WA , EK , 58 km north-northwest of Nicholson H /S, disused racecourse near Linnekar Yard on Linnekar Ck , under debris of old structures, woodland, 17°32 ′ 28 ′′ S , 128°42 ′ 37 ′′ E (coll. R. Crookshanks , 29.i.2005 ) ( AM C.475765). Paratypes : AUSTRALIA , WA , EK , same data as for holotype, 11 dry, four wet ( AM C.443299); Halls Creek District , isolated limestone area south-east of Flora Valley , limestone gully with patches of vine thicket, open slopes, under spinifex, in talus, 18°18 ′ 11 ′′ S , 128°07 ′ 08 ′′ E (coll. V. Kessner , R. Hokkanen, 1.ix.2009 ; WA-74 /09), 51 dry, seven wet ( WAM S49204 ); Springvale-Lansdowne Rd, Little Gold River drainage, 93 km north-west of Halls Creek , on and in soil between rocks of roadside scree under palm, 17°31 ′ 10 ′′ S , 127°11 ′ 52 ′′ E (coll. R. Crookshanks , 31.i.2005 ), five dry, two wet ( AM C.443294) . Other material: AUSTRALIA , WA , EK , Halls Creek District , isolated limestone area south-east of Flora Valley , limestone gully with patches of vine thicket, open slopes, under spinifex, in talus, 18°18 ′ 11 ′′ S , 128°07 ′ 08 ′′ E (coll. V. Kessner , R. Hokkanen, 1.ix.2009 ), 51 dry ( WA-74 /09, WAM S49136 ) . Etymology: In reference to the type locality, Latinized and derived from ‘flora’ and ‘vallis’ (Latin = valley), nouns of female gender in genitive case. Description: Shell ( Figs 4E , 5D , 9D–F ). Moderately large, moderately elevated, thin; micropustulations present; background colour brownish horn fading to whitish towards shell base, with one conspicuous, continuous, relatively wide, brown-reddish peripheral band; no maculations. Genitalia ( Fig. 11 ). Epiphallus as long as penis. Vas deferens entering penial sheath halfway up. Penis four times longer than wide, not coiled inside sheath, with inner wall supporting several irregularly shaped, partly interrupted or corrugated longitudinal pilasters, tapering in prominence posteriorly. Vagina about half as long as penis and two times shorter than free oviduct. Bursa copulatrix with long, pointed well-differentiated end. Remarks: Micropustulation much finer than in E. rivifontis . Shell colour pattern very similar to E. calciphila , E. gregoriana , E. limbunya , E. rivifontis , and E. unistriata . Penis length relative to width (pl/pw) smaller than average and E. rivifontis ; epiphallus length relative to penis (el/pl) 1.5 times the average and as long as in E. rivifontis . Penial inner wall sculpture very similar to E. rivifontis . Vagina length relative to penis (vl/pl) average and larger than in E. rivifontis , vagina length relative to free oviduct (vl/ol) larger than average and E. rivifontis .