Identity of some recently described Lepidoptera from France-re-assessed with DNA barcodes and morphology Author Huemer, Peter Tiroler Landesmuseen Betriebsges. m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall, Austria Author Van Nieukerken, Erik J. 0000-0002-5721-1840 Naturalis Biodiversity Center, PO Box 9517, NL- 2300 RA Leiden, Netherlands. https: // orcid. org / 0000 - 0002 - 5721 - 1840 Corresponding author. p. huemer @ tiroler-landesmuseen. at; https: // orcid. org / 0000 - 0002 - 0630 - 545 X p.huemer@tiroler-landesmuseen.at text Zootaxa 2021 2021-03-09 4941 3 301 337 journal article 10.11646/zootaxa.4941.3.1 1175-5326 4595123 EAFCA1C4-7C08-487F-BA6E-DD74D0A33E19 Cydia oxytropidana Nel & Varenne, 2016 Cydia oxytropidana Nel & Varenne 2016a: 36 , figs. 18, 19a, 20a. Examined material. Holotype Ƌ: France , Hautes-Alpes , La Roche-de-Rame , 30.V.2015 , J. Nel leg. ( TLMF ) . Paratype Ƌ, idem ( TLMF ) . Paratype ♀ , idem ( TLMF ) . Holotype : BOLD Specimen ID TLMF 21933 ; BOLD Sequence ID LEASS475-17 . Paratype : BOLD Specimen ID TLMF 22248 ; BOLD Sequence ID LEASS600-17 . Paratype : BOLD Specimen ID TLMF 22249 ; BOLD Sequence ID LEASS601-17 . BIN: BOLD:ACZ9660 Identity: Cydia oxytropidis (Martini, 1912) , syn. nov. ( Fig. 6 ) FIGURE 6 . Neighbor-joining tree of Cydia oxytropidis incl. C . oxytropidana and the closest European species C . medicaginis (Kuznetsov, 1962) in BOLD. Remarks. Cydia oxytropidana was described from a male holotype and several paratypes of both sexes. Nel & Varenne (2016a) compared the species with C. oxytropidis (Martini, 1912) but based their diagnosis exclusively on genitalia figures published by Razowski (2003) . Whereas the discussed differences in male genitalia are subtle, the females of both species seemed to differ strikingly e.g. by the lack of signa in C. oxytropidis vs well developed in C. oxytropidana . However, re-assessment of allegedly diagnostic characters gave a different picture. Male genitalia show intraspecific variation that does not support two species. Similarly female genitalia characters underly some variation as shown by Nel & Varenne (2016a) for C. oxytropidana . Paired signa in female genitalia are a basic structure in Cydia and well developed in all European species. It is therefore suspected that the figure of C. oxytropidis in Razowski (2003) , based on a specimen from Ukraine , is erroneous. Other specimens of C. oxytropidis from the Ukraine have normally developed signa ( Kavurka 2010 ) and the same applies to material from Central Europe dissected by PH . Finally we have been able to obtain DNA barcode sequences of the holotype and two paratypes of C. oxytropidana which also fully correspond to C. oxytropidis . We therefore synonymize C. oxytropidana with C. oxytropidis .