Order Rodentia - Family Sciuridae Author Robert S. Hoffmann Author Charles G. Anderson text 1993 Smithsonian Institution Press Washington and London Editor Don E. Wilson Editor DeeAnn M. Reeder Mammal Species of the World (2 nd Edition) 419 465 book chapter http://doi.org/10.5281/zenodo.7353130 1-56098-217-9 7353130 Spermophilopsis leptodactylus (Lichtenstein, 1823) . Naturh. Abh. Eversmann's Reise, p. 119 . TYPE LOCALITY: "Vicinity of Kara Ata, 140 km northwest of the Old Town of Bukhara" [ Uzbekistan ] ( Ognev, 1966:394 ) . DISTRIBUTION: SE Kazakstan, Turkmenistan , Uzbekistan , W Tadzhikistan , NE Iran , NW Afghanistan . SYNONYMS: bactrianus (Scully, 1888); heptopotamicus Heptner and Ismagilov, 1952; schumakovi (Satunin, 1908); turcomanus (Eichwald, 1834). COMMENTS: Related to African xerine squirrels ( Atlantoxerus and Xerus ) ( Nadler et al., 1969 ; Nadler and Hoffmann, 1974 ).