A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia Author Zonstein, Sergei Author Kunt, Kadir B. Author Yağmur, Ersen A. text European Journal of Taxonomy 2018 2018-01-30 399 1 93 journal article 10.5852/ejt.2018.399 b389e1f3-82be-4258-9b09-b7ef9d839f99 2118-9773 1163422 E836E138-D6E2-4F62-B4B3-CE2E073F2B24 Raveniola vonwicki Zonstein, 2000 Figs 64, 67 , 82 , 94 , 115 , 127 , 143 , 163 , 195–196 , 210 , 222 Raveniola vonwicki Zonstein, 2000 : 49 , figs 1–3 (♂♀). Diagnosis In the shape of the bulb this species resembles R. marusiki sp. nov. and R. zaitzevi . It differs from both species by a noticeably longer apical segment of the PLS ( Fig. 67 ; cf. Fig. 66 ). The males can be distinguished in having a slightly longer embolus lacking dentate keels ( Figs 195–196 ; cf. Figs 191– 194 ) and a more slender palpal tibia ( Fig. 163 ; cf. Figs 161–162 ). The female of R. vonwicki differs from those in the same species group in having longer spermathecae with stalked, not sessile, lateral receptacles ( Fig. 210 ; cf. Figs 208–209, 211–212 ). Material examined Holotype IRAN : ♂ , Gilan Province , Hishtpar District , E slope of Bogrovdagh Mts , 7 Apr. 1914 , N.S. von Wick leg. ( ZISP 454–914 ). Paratype IRAN : 1 ♀ , collected together with the holotype. Redescription Male (holotype) MEASUREMENTS. TBL 12.00, CL 5.53, CW 4.15, LL 0.43, LW 0.87, SL 2.53, SW 2.27. COLOUR. Carapace, palps and legs dorsally dark brownish rufous; chelicerae reddish brown; sternum, labium, maxillae, palps and legs ventrally light brown; abdomen dorsally brown, with dense light brown spots; ventral surface of abdomen brownish grey. PROSOMA. Carapace and chelicerae as shown in Fig. 82 . Clypeus and eye group as in Fig. 115 . Eye diameters and interdistances: AME 0.16(0.21), ALE 0.27, PLE 0.13, PME 0.14, AME–AME 0.11(0.06), ALE–AME 0.11(0.09), ALE–PLE 0.09, PLE–PME 0.05, PME–PME 0.35. Each cheliceral furrow with 9–10 promarginal teeth and 4–5 mesobasal denticles. Maxillae with 20–24 cuspules each ( Fig. 64 ). LEGS. Tibia and metatarsus I as shown in Fig. 143 . Scopula: distal on metatarsi I–II, entire on tarsi I–II, vestigial on tarsi III, absent on tarsi IV. Trichobothria: 2 rows of 7–8 each on tibiae, 11–14 on metatarsi, 11–12 on tarsi, 9 on cymbium. Paired tarsal claws: outer margin with 8–9 teeth, 6–9 teeth on inner margin. SPINATION. Palp: femur d1–1–1, pd1, rd1; patella r1; tibia d1–1–1, p1–1–1, v7(8); tarsus d5. Leg I: femur d1–1–b–b, pd1–1–1, rd1(0); tibia p1–1, pv1, rv1–1–1–m–m; metatarsus v2. Leg II: femur d1–1–b–b, pd1–1–1; patella p1; tibia p1–1–1, v2–2–3; metatarsus p1, v1–2–3. Leg III: femur d1–1–b–b, pd1–1–1, rd1–1–1; patella p1, r1; tibia d1–1, p1–1, r1–1–1, v2–2–2; metatarsus d1–1–1, pd1–1–1, p1–1–1, r1– 1–1, v2–2–3. Leg IV: femur d1–1–b–b, pd1–1–1, pr1–1–1; patella p1, r1; tibia d1–1, p1–1–1, r1–1–1, v2–2–3; metatarsus d1–1–1, p1–1–1, r1–1–1, v3–2–2–2. Patella I aspinose. PALP. Tibia, cymbium and palpal organ as shown in Fig. 163 . Embolus consisting of almost straight and slightly dilated proximal part lacking keels and of sharply narrowed, spiraled and flattened apical part ( Figs 195–196 ). SPINNERETS. See Fig. 67 . PMS: length 0.53, diameter 0.17. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.70, 0.63, 0.77; total length 2.10; apical segment digitiform. LEG MEASUREMENTS. ♂(♀) Female (paratype)
Femur Patella Tibia Metatarsus Tarsus Total
Palp 2.75 (3.40) 1.82 (2.23) 2.53 (2.65) – 0.97 (2.17) 8.08 (10.45)
Leg I 4.47 (4.47) 2.85 (3.07) 3.93 (3.05) 3.55 (2.65) 2.20 (2.03) 17.00 (15.27)
Leg II 4.10 (4.05) 2.43 (2.85) 3.30 (3.13) 3.25 (2.70) 2.13 (2.07) 15.21 (14.80)
Leg III 3.65 (3.73) 2.10 (2.15) 2.25 (2.63) 3.77 (3.50) 2.20 (2.07) 13.97 (14.08)
Leg IV 4.57 (4.70) 2.17 (2.63) 3.97 (3.87) 5.85 (4.95) 2.43 (2.27) 17.99 (18.42)
MEASUREMENTS. TBL 16.50, CL 6.45, CW 5.67, LL 0.57, LW 1.10, SL 3.25, SW 2.87. COLOUR. As in male. PROSOMA. Carapace and chelicerae as shown in Fig. 94 . Clypeus and eye group as in Fig. 127 . Eye diameters and interdistances: AME 0.18(0.23), ALE 0.31, PLE 0.20, PME 0.18, AME–AME 0.22(0.17), ALE–AME 0.12(0.09), ALE–PLE 0.13, PLE–PME 0.05, PME–PME 0.54. Each cheliceral furrow with 10 promarginal teeth and 6–8 mesobasal denticles. Maxillae with 22–23 cuspules each. LEGS. Scopula: distal on metatarsi I–II, entire on palpal tarsus and tarsi I–II, absent on tarsi III–IV. Trichobothria: 2 rows of 5–7 each on tibiae, 7–11 on metatarsi, 8–10 on tarsi. Paired tarsal claws: outer margin with 5–6 teeth, 5–7 teeth on inner margin. Palpal claw with 5 promarginal teeth. SPINATION. All femora with one basodorsal spine and 3–4 stiff dorsal bristles (weak spines) located medially and distally; patella I aspinose. Palp: femur pd1; patella p1; tibia v2–2–3; tarsus v3(8). Leg I: femur pd1; tibia p1–1, v2–2–3; metatarsus v1–2–2(2–2–3). Leg II: femur pd1; patella p1; tibia p1–1, v2–2–3; metatarsus v2–2–3. Leg III: femur pd1, rd1–1; patella p1, r1; tibia d1(0), p1–1, r1–1, v2–2–3; metatarsus d1–1, p1–1–1–1, r1–1–1–1, v2–2–2–3. Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p1–1, r1–1, v2–2–3; metatarsus d1–1, p1–1–1–1, r1–1–1–1, v2–2–2–3. SPERMATHECAE. Narrow and relatively long, with stalked lateral receptacles ( Fig. 210 ). SPINNERETS. PMS: length 0.73, diameter 0.25. PLS: maximal diameter 0.70; length of basal, medial and apical segments 1.15, 0.85, 1.10; total length 3.10; apical segment digitiform. Ecology Unknown. Distribution Known only from the type locality.